<?xml version="1.0" encoding="UTF-8"?><!DOCTYPE article PUBLIC "-//NLM//DTD JATS (Z39.96) Journal Publishing DTD v1.3 20210610//EN" "https://jats.nlm.nih.gov/publishing/1.3/JATS-journalpublishing1-3.dtd"><article xml:lang="id" dtd-version="1.3" xmlns:xlink="http://www.w3.org/1999/xlink" xmlns:ali="http://www.niso.org/schemas/ali/1.0/" article-type="research-article"><front><journal-meta><journal-id journal-id-type="issn">2089-0257</journal-id><journal-title-group><journal-title>Jurnal Entomologi Indonesia</journal-title><abbrev-journal-title>J Entomol Indones</abbrev-journal-title></journal-title-group><issn pub-type="epub">2089-0257</issn><issn pub-type="ppub">1829-7722</issn><publisher><publisher-name>Perhimpunan Entomologi Indonesia</publisher-name></publisher></journal-meta><article-meta><article-id pub-id-type="doi">10.5994/jei.23.1.11</article-id><title-group><article-title>Keanekaragaman Diptera kanopi pada empat tipe penggunaan lahan berbeda di Jambi, Sumatra, Indonesia</article-title><subtitle>The diversity of dipteran canopy on the four different land use types in Jambi, Sumatra, Indonesia</subtitle></title-group><contrib-group><contrib contrib-type="author"><contrib-id contrib-id-type="orcid">https://orcid.org/0009-0009-9963-0689</contrib-id><name><surname>Ulinnuha</surname><given-names>Ulfa</given-names></name><address><country>Indonesia</country></address><xref ref-type="aff" rid="AFF-1"></xref></contrib><contrib contrib-type="author"><contrib-id contrib-id-type="orcid">https://orcid.org/0000-0002-2843-0737</contrib-id><name><surname>Buchori</surname><given-names>Damayanti</given-names></name><address><country>Indonesia</country></address><xref ref-type="aff" rid="AFF-2"></xref></contrib><contrib contrib-type="author"><contrib-id contrib-id-type="orcid">https://orcid.org/0000-0002-5162-9779</contrib-id><name><surname>Drescher</surname><given-names>Jochen</given-names></name><address><country>Germany</country></address><xref ref-type="aff" rid="AFF-3"></xref></contrib><contrib contrib-type="author"><contrib-id contrib-id-type="orcid">https://orcid.org/0000-0003-4350-9520</contrib-id><name><surname>Scheu</surname><given-names>Stefan</given-names></name><address><country>Germany</country></address><xref ref-type="aff" rid="AFF-3"></xref></contrib><contrib contrib-type="author"><contrib-id contrib-id-type="orcid">https://orcid.org/0000-0003-4222-411X</contrib-id><name><surname>Nazarreta</surname><given-names>Rizky</given-names></name><address><country>Indonesia</country></address><xref rid="AFF-4" ref-type="aff"></xref></contrib><contrib contrib-type="author"><contrib-id contrib-id-type="orcid">https://orcid.org/0000-0003-1227-2794</contrib-id><name><surname>Kasmiatun</surname></name><address><country>Indonesia</country></address><xref ref-type="aff" rid="AFF-5"></xref></contrib><contrib contrib-type="author"><name><surname>Widhyastuti</surname><given-names>Paramita Hayu</given-names></name><address><country>Indonesia</country></address><xref ref-type="aff" rid="AFF-6"></xref></contrib><contrib contrib-type="author"><contrib-id contrib-id-type="orcid">https://orcid.org/0000-0002-6673-6198</contrib-id><name><surname>Rizali</surname><given-names>Akhmad</given-names></name><address><country>Indonesia</country></address><xref ref-type="aff" rid="AFF-6"></xref></contrib><contrib contrib-type="author"><contrib-id contrib-id-type="orcid">https://orcid.org/0000-0001-9507-6275</contrib-id><name><surname>Hidayat</surname><given-names>Purnama</given-names></name><address><country>Indonesia</country><email>phidayat@apps.ipb.ac.id</email></address><xref ref-type="aff" rid="AFF-4"></xref><xref ref-type="corresp" rid="cor-8"></xref></contrib></contrib-group><contrib-group><contrib contrib-type="editor"><name><surname>Shahabuddin</surname><given-names>Shahabuddin</given-names></name><address><country>Indonesia</country></address><xref ref-type="aff" rid="EDITOR-AFF-1"></xref></contrib></contrib-group><aff id="AFF-1"><institution-wrap><institution>Bogor Agricultural University</institution><institution-id institution-id-type="ror">https://ror.org/05smgpd89</institution-id></institution-wrap><country country="ID">Indonesia</country></aff><aff id="AFF-2"><institution content-type="dept">Departemen Proteksi Tanaman, Fakultas Pertanian</institution><institution-wrap><institution>IPB University</institution><institution-id institution-id-type="ror">https://ror.org/05smgpd89</institution-id></institution-wrap><country country="ID">Indonesia</country></aff><aff id="AFF-3"><institution content-type="dept">Department of Animal Ecology, J. F. Blumenbach Institute of Zoology and Anthropology</institution><institution-wrap><institution>University of Göttingen</institution><institution-id institution-id-type="ror">https://ror.org/01y9bpm73</institution-id></institution-wrap><addr-line>Göttingen 37073</addr-line><country country="DE">Germany</country></aff><aff id="AFF-4"><institution content-type="dept">Departemen Proteksi Tanaman, Fakultas Pertanian</institution><institution-wrap><institution>IPB University</institution><institution-id institution-id-type="ror">https://ror.org/05smgpd89</institution-id></institution-wrap><addr-line>Jalan Kamper, Kampus IPB Dramaga, Bogor 16680</addr-line><country country="ID">Indonesia</country></aff><aff id="AFF-5">Pusat Penelitian Kelapa Sawit (PPKS), Divisi Proteksi Tanaman, Jalan Pematangsiantar - Tanah Jawa, Marihat, Sumatera Utara 21128, Indonesia</aff><aff id="AFF-6"><institution content-type="dept">Jurusan Hama dan Penyakit Tumbuhan, Fakultas Pertanian</institution><institution-wrap><institution>Universitas Brawijaya</institution><institution-id institution-id-type="ror">https://ror.org/01wk3d929</institution-id></institution-wrap><addr-line>Jalan Veteran, Malang 65145</addr-line><country country="ID">Indonesia</country></aff><aff id="EDITOR-AFF-1"><institution content-type="dept">Program Studi Agroteknologi, Fakultas Pertanian</institution><institution-wrap><institution>Universitas Tadulako</institution><institution-id institution-id-type="ror">https://ror.org/01z0mc198</institution-id></institution-wrap><country country="ID">Palu</country></aff><author-notes><corresp id="cor-8">Corresponding author: Purnama Hidayat, Departemen Proteksi Tanaman, Fakultas Pertanian, IPB University, Jalan Kamper, Kampus IPB Dramaga, Bogor 16680, Indonesia.  Email: <email>phidayat@apps.ipb.ac.id</email></corresp></author-notes><pub-date date-type="pub" iso-8601-date="2026-4-24" publication-format="electronic"><day>24</day><month>4</month><year>2026</year></pub-date><pub-date iso-8601-date="2026-4-12" publication-format="electronic" date-type="collection"><day>12</day><month>4</month><year>2026</year></pub-date><volume>23</volume><issue>1</issue><issue-title>Vol. 23 No. 1 (2026): March</issue-title><fpage>11</fpage><lpage>29</lpage><history><date date-type="received" iso-8601-date="2024-7-5"><day>5</day><month>7</month><year>2024</year></date><date iso-8601-date="2026-4-23" date-type="accepted"><day>23</day><month>4</month><year>2026</year></date></history><permissions><copyright-statement>Hak Cipta (c) 2026</copyright-statement><copyright-year>2026</copyright-year><copyright-holder>The Author(s).</copyright-holder><license license-type="open-access" xlink:href="https://creativecommons.org/licenses/by/4.0/"><ali:license_ref xmlns:ali="http://www.niso.org/schemas/ali/1.0/">https://creativecommons.org/licenses/by/4.0/</ali:license_ref><license-p>Artikel ini berlisensi  Creative Commons Attribution 4.0 International License.</license-p></license></permissions><self-uri xlink:href="https://jurnal.pei-pusat.org/index.php/jei/article/view/890" xlink:title="Keanekaragaman Diptera kanopi pada empat tipe penggunaan lahan berbeda di Jambi, Sumatra, Indonesia">Keanekaragaman Diptera kanopi pada empat tipe penggunaan lahan berbeda di Jambi, Sumatra, Indonesia</self-uri><abstract><p>Alih fungsi hutan menjadi perkebunan monokultur dapat menyebabkan penurunan tutupan kanopi pohon dan keanekaragaman serangga, salah satunya Diptera. Diptera merupakan kelompok serangga yang berperan penting bagi keseimbangan ekosistem. Penelitian ini bertujuan untuk memelajari dampak konversi hutan menjadi perkebunan monokultur terhadap keanekaragaman dan komposisi Diptera pada empat tipe penggunaan lahan di Jambi, Sumatra. Pengambilan sampel Diptera dilakukan dengan menggunakan metode pengasapan kanopi pohon pada dua lanskap, yaitu di dalam dan sekitar area Taman Nasional Bukit Duabelas dan Hutan Harapan. Pada masing-masing lanskap, ditentukan empat tipe penggunaan lahan, yaitu hutan alami, hutan karet, perkebunan karet, dan perkebunan kelapa sawit. Total individu Diptera yang dikoleksi sejumlah 13.513 individu, terdiri atas 39 famili, 169 genus, dan 399 spesies. Adapun famili yang paling dominan yaitu Cecidomyiidae, sementara spesies yang paling dominan yaitu <italic>Procontarinia </italic>sp.01. Kelimpahan dan kekayaan spesies Diptera tertinggi ditemukan di hutan alami, kemudian diikuti hutan karet. Kelimpahan terendah ditemukan di perkebunan kelapa sawit, sedangkan kekayaan spesies terendah ditemukan di perkebunan karet. Hasil uji ANOVA menunjukkan tipe penggunaan lahan berbeda berpengaruh terhadap kelimpahan dan kekayaan spesies Diptera. Sementara itu, perbedaan lanskap tidak berpengaruh terhadap kelimpahan dan kekayaan spesies Diptera. Komposisi spesies Diptera dipengaruhi oleh tipe penggunaan lahan, lanskap, dan interaksi keduanya. Komposisi Diptera di hutan alami dan hutan karet menunjukkan kemiripan tertinggi, sedangkan kemiripan terendah pada hutan alami dan perkebunan kelapa sawit. Perubahan komposisi ini didorong oleh spesies <italic>turnover.</italic> Selain itu, konversi lahan juga menyebabkan peralihan dominansi, hutan didominasi oleh famili Cecidomyiidae, lalu dominasi beralih ke famili Ceratopogonidae di perkebunan monokultur.</p></abstract><kwd-group><kwd>lalat</kwd><kwd>nyamuk</kwd><kwd>perubahan tata guna lahan</kwd><kwd>serangga arboreal</kwd><kwd>sawit</kwd><kwd>karet</kwd></kwd-group><custom-meta-group><custom-meta><meta-name>File created by JATS Editor</meta-name><meta-value><ext-link ext-link-type="uri" xlink:href="https://jatseditor.com" xlink:title="JATS Editor">JATS Editor</ext-link></meta-value></custom-meta><custom-meta><meta-name>issue-created-year</meta-name><meta-value>2026</meta-value></custom-meta></custom-meta-group></article-meta></front><body><sec><title>PENDAHULUAN</title><p>Hutan hujan tropis dataran rendah merupakan kawasan dengan ekosistem yang kaya akan keanekaragaman hayati flora dan fauna<xref ref-type="bibr" rid="BIBR-45">(Myers et al., 2000)</xref>, akan tetapi keberlangsungannya terancam akibat meningkatnya laju konversi hutan<xref ref-type="bibr" rid="BIBR-26">(Gibson et al., 2011)</xref>. Perubahan fungsi hutan menjadi lahan perkebunan, khususnya untuk karet dan kelapa sawit menjadi salah satu faktor utama penurunan luas area hutan primer di Jambi<xref ref-type="bibr" rid="BIBR-18">(Drescher et al., 2016)</xref>. Alih fungsi hutan tersebut tidak hanya menurunkan jumlah populasi dan spesies yang hidup di dalamnya, akan tetapi juga berpengaruh signifikan pada proses distribusi energi, khususnya antara jejaring makanan di atas permukaan tanah (herbivor dan predator) dan di bawah tanah (fungivor, bakterivor, dan dekomposer)<xref ref-type="bibr" rid="BIBR-53">(Potapov et al., 2024)</xref>. Berdasarkan laporan Margono et al. (2014)<xref rid="BIBR-41" ref-type="bibr">(Margono et al., 2014)</xref> terdapat beberapa pulau di Indonesia yang terus mengalami alih fungsi hutan menjadi kawasan perkebunan, di antaranya Sumatra, Kalimantan, dan Papua<xref ref-type="bibr" rid="BIBR-41">(Margono et al., 2014)</xref>. Salah satu provinsi di pulau Sumatra yang mengalami alih fungsi hutan menjadi kawasan perkebunan karet dan kelapa sawit, yaitu Provinsi Jambi. Luas area hutan di Jambi terus mengalami penurunan sejak tahun 1970-an sehingga hanya tersisa sebesar 28% pada tahun 2023<xref ref-type="bibr" rid="BIBR-39">(Laumonier et al., 2010)</xref>;<xref ref-type="bibr" rid="BIBR-9">(Jambi, 2025)</xref>. Adapun luas area perkebunan kelapa sawit dan karet saat ini dilaporkan berturut-turut sebesar 1.135.357 hektar atau 60% dan 601.754 hektar atau 32% dari seluruh perkebunan di Jambi<xref ref-type="bibr" rid="BIBR-10">(Jambi, 2025)</xref>.</p><p>Konversi hutan menjadi kawasan perkebunan monokultur diketahui berdampak negatif bagi berbagai spesies yang hidup di dalamnya, di antaranya dapat menurunkan keanekaragaman spesies tumbuhan, khususnya yang bergantung pada ekosistem hutan primer<xref ref-type="bibr" rid="BIBR-59">(Rembold et al., 2017)</xref>. Selain itu, diketahui juga dapat meningkatkan ancaman spesies invasif yang dapat mengganggu kehidupan spesies asli di kawasan hutan tersebut<xref rid="BIBR-59" ref-type="bibr">(Rembold et al., 2017)</xref>. Lebih lanjut, alih fungsi lahan juga berpengaruh signifikan terhadap keberlanjutan hidup berbagai fauna, khususnya serangga. Beberapa penelitian di Jambi melaporkan adanya penurunan kelimpahan dan kekayaan berbagai kelompok serangga akibat adanya alih fungsi hutan, antara lain semut<xref ref-type="bibr" rid="BIBR-47">(Nazarreta et al., 2020)</xref>;<xref ref-type="bibr" rid="BIBR-17">(Desriana et al., 2023)</xref>;<xref rid="BIBR-19" ref-type="bibr">(TR et al., 2024)</xref>, kupu-kupu<xref ref-type="bibr" rid="BIBR-50">(Panjaitan et al., 2020)</xref>, Hymenoptera parasitoid<xref ref-type="bibr" rid="BIBR-5">(Azhar et al., 2022)</xref>, kumbang Curculionidae, Chrysomelidae, Elateridae, dan Staphylinidae<xref ref-type="bibr" rid="BIBR-34">(Kasmiatun et al., 2022)</xref>. Penurunan keanekaragaman serangga ini menyebabkan perubahan aliran energi antartrofik dalam jejaring makanan dan menurunkan fungsi ekologisnya<xref ref-type="bibr" rid="BIBR-52">(Pollierer et al., 2023)</xref>.</p><p>Penelitian mengenai dampak perubahan tata guna lahan pada berbagai jenis serangga saat ini semakin berkembang dan telah banyak dilaporkan, akan tetapi belum banyak yang melaporkan secara khusus terkait Diptera. Ordo Diptera merupakan kelompok serangga yang memiliki ciri morfologi berupa sepasang sayap dan sepasang halter, terbagi menjadi dua subordo, yaitu Brachycera dan Nematocera<xref ref-type="bibr" rid="BIBR-36">(Kirk-Spriggs et al., 2017)</xref>. Diptera mempunyai peran yang penting di alam, yaitu sebagai musuh alami (parasitoid, predator), herbivor (fitofag), penyerbuk, dan pengurai<xref ref-type="bibr" rid="BIBR-36">(Kirk-Spriggs et al., 2017)</xref>. Sejauh ini, telah dilaporkan bahwa terdapat 176.080 spesies Diptera yang telah dideskripsikan dari seluruh dunia, terdiri atas 17.973 genus dan 298 famili<xref ref-type="bibr" rid="BIBR-23">(G.B.I.F., 2025)</xref>. Diptera memiliki penyebaran yang luas dan dapat hidup di berbagai tipe habitat, seperti di peternakan<xref ref-type="bibr" rid="BIBR-73">(Zimmer et al., 2014)</xref>, pemukiman<xref ref-type="bibr" rid="BIBR-14">(Chen et al., 2016)</xref>, pegunungan<xref ref-type="bibr" rid="BIBR-46">(Navai et al., 2017)</xref>, hutan<xref ref-type="bibr" rid="BIBR-56">(Quaglia et al., 2020)</xref>, sekitar danau<xref ref-type="bibr" rid="BIBR-20">(Erram &amp; Burkett-Cadena, 2021)</xref>, dan habitat dengan tumbuhan berbunga<xref ref-type="bibr" rid="BIBR-30">(Heiduk et al., 2023)</xref>. Lanskap dengan hutan yang luas diketahui memiliki kelimpahan dan kekayaan spesies Diptera yang lebih tinggi dibandingkan dengan lanskap dengan hutan yang terfragmentasi <xref ref-type="bibr" rid="BIBR-62">(Smith &amp; Mayfield, 2015)</xref>. Keberadaan Diptera pada suatu ekosistem dipengaruhi oleh komposisi dan struktur vegetasi, juga iklim mikro<xref ref-type="bibr" rid="BIBR-36">(Kirk-Spriggs et al., 2017)</xref>. Tutupan kanopi menjadi salah satu faktor penting pendukung keberlanjutan ekosistem hutan karena dapat memengaruhi iklim mikro di dalam habitat serangga<xref ref-type="bibr" rid="BIBR-18">(Drescher et al., 2016)</xref>. Hasil penelitian Scherber et al. (2014)<xref ref-type="bibr" rid="BIBR-60">(Scherber et al., 2014)</xref>, kelimpahan Diptera mencapai 300 individu/ plot di kawasan dengan tutupan kanopi &gt;90%, sementara kelimpahannya meningkat menjadi 500 individu/plot pada tutupan kanopi 70% sehingga ada kenaikan kelimpahan Diptera ketika tutupan kanopi berkurang. Namun demikian, pola tersebut berbeda dengan hasil penelitian terhadap ordo serangga lain di Jambi, yang telah disebutkan sebelumnya<xref ref-type="bibr" rid="BIBR-47">(Nazarreta et al., 2020)</xref>;<xref ref-type="bibr" rid="BIBR-50">(Panjaitan et al., 2020)</xref>;<xref ref-type="bibr" rid="BIBR-5">(Azhar et al., 2022)</xref>;<xref ref-type="bibr" rid="BIBR-34">(Kasmiatun et al., 2022)</xref>;<xref ref-type="bibr" rid="BIBR-19">(TR et al., 2024)</xref>.</p><p>Penelitian ini dilakukan untuk mengkaji respons Diptera kanopi di Jambi terhadap perubahan tutupan kanopi, dengan membandingkan pola yang dilaporkan oleh Scherber et al. (2014)<xref rid="BIBR-60" ref-type="bibr">(Scherber et al., 2014)</xref> dengan pola yang telah dilaporkan pada ordo serangga lain di Jambi. Penelitian ini secara khusus dilakukan untuk mengetahui dampak alih fungsi lahan hutan terhadap keberadaan Diptera, dengan merujuk penelitian sebelumnya<xref ref-type="bibr" rid="BIBR-52">(Pollierer et al., 2023)</xref>;<xref ref-type="bibr" rid="BIBR-19">(TR et al., 2024)</xref>;<xref ref-type="bibr" rid="BIBR-53">(Potapov et al., 2024)</xref>. Adapun tujuan dari penelitian ini, yaitu untuk memelajari pengaruh perubahan tipe penggunaan lahan hutan menjadi kawasan agroforestri dan perkebunan monokultur terhadap keanekaragaman dan komposisi Diptera kanopi. Hasil penelitian diharapkan dapat memberikan data awal mengenai keanekaragaman dan komposisi serangga Diptera kanopi di hutan alami, hutan karet, perkebunan karet, dan perkebunan kelapa sawit, di Jambi, Sumatra.</p></sec><sec><title>BAHAN DAN METODE</title><sec><title>Waktu dan lokasi penelitian</title><p>Penelitian ini merupakan bagian dari kerja sama penelitian jangka panjang antara Indonesia dan Jerman, CRC990/EFForTS (<italic>Collaborative Research Centre 990: Ecological and Socioeconomic Functions of Tropical Lowland Rainforest Transformation Systems</italic>) Fase 1, dengan nomor izin koleksi sampel S.710/KKH-2/2013 yang diterbitkan oleh Direktorat Jenderal Perlindungan Hutan dan Konservasi Alam (PHKA), Kementerian Kehutanan Indonesia. Proses koleksi sampel dilakukan dari bulan Juli hingga Oktober 2013 pada musim kemarau, di dalam dan sekitar kawasan Taman Nasional Bukit Duabelas dan Hutan Harapan, Jambi, Sumatra. Taman Nasional Bukit Duabelas merupakan kawasan konservasi yang dikelola oleh Balai Konservasi Sumberdaya Alam Jambi. Sementara itu, kawasan Hutan Harapan merupakan area hutan sekunder yang telah dilakukan upaya restorasi oleh PT. REKI (Restorasi Ekosistem Konservasi Indonesia). Adapun proses identifikasi sampel yang telah terkoleksi dilakukan dari bulan Februari hingga Juli 2018 dan Desember 2018 hingga April 2019 di Laboratorium Pengendalian Hayati, Departemen Proteksi Tanaman, Fakultas Pertanian, Institut Pertanian Bogor.</p></sec><sec><title>Plot pengamatan</title><p>Kawasan di dalam dan sekitar Taman Nasional Bukit Duabelas dan Hutan Harapan dikategorikan sebagai skala lanskap. Pada masing-masing lanskap terdapat empat tipe penggunaan lahan, yaitu hutan alami, hutan karet, perkebunan karet, dan perkebunan kelapa sawit. Tipe penggunaan lahan hutan alami merupakan kawasan hutan primer yang terdegradasi dan mengalami penebangan selektif, akan tetapi tidak banyak mengalami gangguan dari manusia<xref rid="BIBR-41" ref-type="bibr">(Margono et al., 2014)</xref>. Sementara itu, hutan karet merupakan kawasan agroforestri dengan sistem pertanian ekstensif, yang dikembangkan melalui penanaman tanaman karet di kawasan hutan sekunder atau hutan yang sebelumnya mengalami kerusakan, kemudian dibiarkan tumbuh secara alami tanpa perawatan intensif<xref ref-type="bibr" rid="BIBR-59">(Rembold et al., 2017)</xref>. Adapun kawasan perkebunan karet (<italic>Hevea brasiliensis</italic>) dan perkebunan kelapa sawit (<italic>Elaeis guineensis</italic>) merupakan area perkebunan monokultur yang dikelola masyarakat lokal dengan sistem pertanian intensif<xref ref-type="bibr" rid="BIBR-18">(Drescher et al., 2016)</xref>. Setiap tipe penggunaan lahan ditentukan empat plot inti berukuran 50 m × 50 m dengan jarak antarplot sekitar 40 m (32 plot inti)<xref ref-type="bibr" rid="BIBR-18">(Drescher et al., 2016)</xref>. Proses pengambilan sampel dilakukan sebanyak tiga ulangan pada kanopi pohon berbeda dengan luas area sekitar 5 m × 5 m yang kemudian dijadikan sebagai subplot. Total subplot yang digunakan sebanyak 96 subplot (2 lanskap × 4 tipe penggunaan lahan × 4 plot inti × 3 ulangan) (<xref ref-type="fig" rid="figure-1">Gambar 1</xref>).</p><fig id="figure-1" ignoredToc=""><label>Gambar 1</label><caption><p>Lokasi penelitian EFForTS di Jambi<xref ref-type="bibr" rid="BIBR-3">(Amrulloh et al., 2022)</xref>.</p><p>Location of EFForTS study sites in Jambi<xref ref-type="bibr" rid="BIBR-3">(Amrulloh et al., 2022)</xref></p></caption><graphic xlink:href="https://jurnal.pei-pusat.org/index.php/jei/article/download/890/655/9028" mime-subtype="png" mimetype="image"><alt-text>Image</alt-text></graphic></fig></sec><sec><title>Pengambilan sampel</title><p>Pengambilan sampel Diptera kanopi dilakukan dengan menggunakan metode pengasapan kanopi (<italic>fogging</italic>). Adapun pohon yang dipilih untuk proses pengasapan merupakan pohon dengan kanopi yang mewakili kondisi habitat pada area plot tersebut. Pengasapan kanopi pohon dilakukan dengan menggunakan alat Swingfog® SN50 <italic>fogger</italic> (Swingtec GmbH, Germany) dengan bahan yang dilarutkan berupa 50 ml DECIS 25 EC® (insektisida piretroid dengan bahan aktif deltametrin, 25 g/l) dan 4,0–4,5 l minyak putih petroleum. Insektisida piretroid bersifat <italic>knockdown</italic> yang dapat melemahkan sistem motorik atau menyebabkan kelumpuhan sementara pada serangga<xref ref-type="bibr" rid="BIBR-4">(Athanassiou et al., 2021)</xref>. Proses pengasapan dilakukan pada pagi hari sekitar pukul 06.00 WIB dengan mengarahkan alat <italic>fogging</italic> ke kanopi pohon target selama 20 menit. Sebanyak 16 wadah penampung berbentuk limas terbalik berbahan nilon berukuran 1 m × 1 m dipasang mengelilingi pohon di bawah kanopi target. Pada bagian bawah wadah penampung tersebut juga dipasang botol koleksi berisi alkohol 70% untuk menampung serangga yang mengalami efek <italic>knockdown</italic>. Setelah 2 jam pengasapan, serangga yang jatuh ke wadah penampung tersebut dikumpulkan, dimasukkan ke dalam botol sampel, lalu diberi label sesuai kode plot dan tanggal pengambilan sampel. Botol berisi sampel tersebut dibawa ke laboratorium dan disimpan pada suhu kurang lebih -20°C untuk disortir dan diidentifikasi lebih lanjut.</p></sec><sec><title>Identifikasi Diptera</title><p>Sampel Diptera kanopi terlebih dahulu dipisahkan dari sampel artropoda lainnya. Kemudian sampel tersebut diidentifikasi lebih lanjut berdasarkan tingkat subordo, famili, hingga morfospesies. Proses identifikasi dilakukan dengan menggunakan beberapa buku kunci identifikasi, antara lain McAlpine et al. (1981, 1987)<xref rid="BIBR-42" ref-type="bibr">(McAlpine et al., 1981)</xref>;<xref ref-type="bibr" rid="BIBR-43">(McAlpine et al., 1987)</xref>, Kirk-Spriggs et al. (2017a, 2017b, 2021)<xref ref-type="bibr" rid="BIBR-36">(Kirk-Spriggs et al., 2017)</xref>;<xref ref-type="bibr" rid="BIBR-37">(Kirk-Spriggs et al., 2017)</xref>;<xref ref-type="bibr" rid="BIBR-38">(Kirk-Spriggs et al., 2021)</xref>, dan Nugroho &amp; Mujiyono (2021)<xref ref-type="bibr" rid="BIBR-48">(Nugroho &amp; Mujiyono, 2021)</xref>. Setelah proses identifikasi selesai, sebanyak satu hingga dua individu dari setiap morfospesies dipisahkan dan dijadikan sampel master.</p></sec><sec><title>Analisis data</title><p>Analisis data dilakukan dengan menggunakan perangkat lunak R 4.5.2 dan RStudio 2025.09.2+418<xref ref-type="bibr" rid="BIBR-57">(Team, 2025)</xref>. Kode R yang digunakan merupakan hasil modifikasi dari analisis Junggebauer et al. (2021)<xref ref-type="bibr" rid="BIBR-33">(Junggebauer et al., 2021)</xref>, Ramos et al. (2022)<xref ref-type="bibr" rid="BIBR-58">(Ramos et al., 2022)</xref>, dan Ehlers et al. (2024)<xref ref-type="bibr" rid="BIBR-19">(TR et al., 2024)</xref>, yang divisualisasikan dengan <italic>ggplot2</italic><xref ref-type="bibr" rid="BIBR-69">(Wickham, 2016)</xref>. Indeks keanekaragaman alfa yang digunakan, yaitu Shannon-Wiener, Simpson, dan Evenness yang dianalisis menggunakan paket <italic>vegan::diversity</italic><xref ref-type="bibr" rid="BIBR-49">(Oksanen et al., 2025)</xref>. Pengaruh perbedaan tipe penggunaan lahan terhadap kelimpahan dan kekayaan spesies Diptera kanopi dianalisis menggunakan analisis sidik ragam (ANOVA) dengan pemodelan <italic>generalized linear model</italic> (GLM), menggunakan Famili Negative Binomial untuk analisis kelimpahan dan Famili Gaussian untuk kekayaan spesies. Famili dipilih berdasarkan visualisasi Q-Q plot dan nilai <italic>akaike information criterion</italic> (AIC) terendah. Uji lanjut menggunakan Tukey dengan taraf α = 5%. Data divisualisasikan menggunakan gabungan <italic>barplot.</italic> Estimasi kekayaan spesies Diptera kanopi dihitung berdasarkan nilai <italic>abundance-based coverage estimator</italic> (ACE) menggunakan perangkat lunak EstimateS 9.10, lalu divisualisasikan menggunakan kurva akumulasi spesies. Peringkat kelimpahan spesies dianalisis menggunakan paket RankAbund<xref ref-type="bibr" rid="BIBR-29">(Hartke, 2020)</xref> untuk mengetahui pola dominansi pada setiap tipe penggunaan lahan. Tumpang tindih spesies dan famili pada empat tipe penggunaan lahan dianalisis menggunakan <italic>vegan</italic>, lalu divisualisasikan menggunakan diagram Venn. Analisis komposisi spesies dilakukan dengan menggunakan model glm multivariat, lalu divisualisasikan dengan boral<xref ref-type="bibr" rid="BIBR-31">(Hui, 2025)</xref> dan ggboral<xref rid="BIBR-12" ref-type="bibr">(Bedward, 2020)</xref>. Penelitian ini juga menganalisis spesies <italic>turnover</italic> yang merupakan bagian dari keanekaragaman beta menggunakan indeks Bray-Curtis. Spesies <italic>turnover</italic> adalah pergantian spesies antar komunitas, ketika ada spesies tertentu yang hilang pada suatu lokasi lalu digantikan oleh spesies yang lain<xref ref-type="bibr" rid="BIBR-11">(Baselga, 2010)</xref>, merupakan konsekuensi dari perubahan kondisi lingkungan<xref ref-type="bibr" rid="BIBR-55">(Qian et al., 2005)</xref>.</p></sec></sec><sec><title>HASIL</title><p>Total Diptera kanopi yang ditemukan sejumlah 13.513 individu, terdiri atas 39 famili, 73 subfamili, 169 genus, dan 399 spesies. Subordo Nematocera memiliki kelimpahan yang lebih tinggi dibandingkan dengan Subordo Brachycera, yaitu 11.329 dan 2.184 individu (<xref rid="table-1" ref-type="table">Tabel 1</xref>). Adapun famili yang mendominasi di seluruh tipe penggunaan lahan, yaitu Cecidomyiidae dari Subordo Nematocera. Sementara itu, lima spesies yang mendominasi hanya mencakup 30,6% dari total individu yang ditemukan, yaitu Procontarinia sp.01 (1.242 individu), Mycodiplosis sp.02 (900 individu), Culicoides sp.01 (789 individu), Miastor sp.02 (729 individu), dan Macrolabis sp.01 (470 individu) (<xref ref-type="table" rid="table-2">Tabel 2</xref>). Berdasarkan masing-masing tipe penggunaan lahan, spesies yang mendominasi di hutan alami dan hutan karet, yaitu Procontarinia sp.01, masing-masing sebanyak 742 dan 390 individu. Kemudian, di perkebunan karet terdapat Brachypogon sp.01 sebanyak 216 individu dan di perkebunan kelapa sawit terdapat Dasyhelea sp.01 sebanyak 103 individu. Adapun persentase spesies Diptera yang berhasil didapatkan dari 32 plot menggunakan metode pengasapan kanopi pohon adalah sebesar 87,4%. Berdasarkan nilai ACE, diperkirakan masih terdapat spesies yang belum terdeteksi pada setiap tipe penggunaan lahan (<xref ref-type="table" rid="table-3">Tabel 3</xref>). Kemiringan kurva akumulasi spesies yang masih curam mengindikasikan jumlah spesies Diptera terus bertambah seiring penambahan plot (<xref rid="figure-2" ref-type="fig">Gambar 2</xref>A).</p><table-wrap id="table-1" ignoredToc=""><label>Tabel 1</label><caption><p>Kelimpahan brachyceran dan nematoceran di empat tipe penggunaan lahan </p><p>Abundance of brachyceran and nematoceran on four types of use</p></caption><table frame="box" rules="all"><thead><tr><th valign="middle" align="left" colspan="1">Subordo</th><th colspan="1" valign="middle" align="center">Hutan alami (<italic>Forest</italic>)</th><th valign="middle" align="center" colspan="1">Hutan karet (<italic>Jungle rubber</italic>)</th><th colspan="1" valign="middle" align="center">Perkebunan karet (<italic>Rubber plantation</italic>)</th><th align="center" colspan="1" valign="middle">Perkebunan kelapa sawit (<italic>Oil palm plantation</italic>)</th><th align="center" colspan="1" valign="middle">Total (<italic>Total</italic>)</th></tr></thead><tbody><tr><td valign="top" align="left" colspan="1">Subordo Brachycera (<italic>Brachycera Suborder</italic>)</td><td valign="top" align="center" colspan="1">783</td><td align="center" colspan="1" valign="top">576</td><td align="center" colspan="1" valign="top">420</td><td valign="top" align="center" colspan="1">405</td><td align="center" colspan="1" valign="top">2.184</td></tr><tr><td colspan="1" valign="top" align="left">Famili (<italic>Family</italic>)</td><td valign="top" align="center" colspan="1">22</td><td valign="top" align="center" colspan="1">21</td><td valign="top" align="center" colspan="1">13</td><td valign="top" align="center" colspan="1">16</td><td align="center" colspan="1" valign="top">27</td></tr><tr><td valign="top" align="left" colspan="1">Subfamili (<italic>Subfamily</italic>)</td><td align="center" colspan="1" valign="top">35</td><td align="center" colspan="1" valign="top">35</td><td valign="top" align="center" colspan="1">25</td><td valign="top" align="center" colspan="1">27</td><td valign="top" align="center" colspan="1">45</td></tr><tr><td valign="top" align="left" colspan="1">Spesies (<italic>Species</italic>)</td><td valign="top" align="center" colspan="1">164</td><td valign="top" align="center" colspan="1">126</td><td align="center" colspan="1" valign="top">71</td><td align="center" colspan="1" valign="top">80</td><td valign="top" align="center" colspan="1">230</td></tr><tr><td valign="top" align="left" colspan="1">Subordo Nematocera (<italic>Nematocera Suborder</italic>)</td><td align="center" colspan="1" valign="top">4.330</td><td colspan="1" valign="top" align="center">3.599</td><td valign="top" align="center" colspan="1">2.144</td><td valign="top" align="center" colspan="1">1.256</td><td valign="top" align="center" colspan="1">11.329</td></tr><tr><td valign="top" align="left" colspan="1">Famili (<italic>Family</italic>)</td><td valign="top" align="center" colspan="1">12</td><td align="center" colspan="1" valign="top">11</td><td valign="top" align="center" colspan="1">10</td><td colspan="1" valign="top" align="center">10</td><td align="center" colspan="1" valign="top">12</td></tr><tr><td valign="top" align="left" colspan="1">Subfamili (<italic>Subfamily</italic>)</td><td valign="top" align="center" colspan="1">24</td><td align="center" colspan="1" valign="top">23</td><td align="center" colspan="1" valign="top">23</td><td align="center" colspan="1" valign="top">23</td><td align="center" colspan="1" valign="top">27</td></tr><tr><td align="left" colspan="1" valign="top">Spesies (<italic>Species</italic>)</td><td valign="top" align="center" colspan="1">123</td><td valign="top" align="center" colspan="1">137</td><td align="center" colspan="1" valign="top">116</td><td valign="top" align="center" colspan="1">121</td><td align="center" colspan="1" valign="top">169</td></tr></tbody></table></table-wrap><table-wrap id="table-2" ignoredToc=""><label>Tabel 2</label><caption><p>Spesies dominan di empat tipe penggunaan lahan</p><p>Dominant species on four land use types</p></caption><table rules="all" frame="box"><thead><tr><th colspan="1" valign="middle" align="left">Tipe penggunaan lahan (<italic>Land use type</italic>)</th><th valign="middle" align="center" colspan="1">Peringkat (<italic>Rank</italic>)</th><th align="center" colspan="1" valign="middle">Spesies (<italic>Species</italic>)</th><th align="center" colspan="1" valign="middle">Famili (<italic>Family</italic>)</th><th align="center" colspan="1" valign="middle">Kelimpahan (<italic>Abundance</italic>)</th><th valign="middle" align="center" colspan="1">%</th></tr></thead><tbody><tr><td align="left" colspan="1" rowspan="6" valign="top">Keseluruhan (<italic>Overall</italic>)</td><td valign="top" align="center" colspan="1">1</td><td valign="top" align="left" colspan="1"><italic>Procontarinia </italic>sp.01</td><td colspan="1" valign="top" align="left">Cecidomyiidae</td><td colspan="1" valign="top" align="center">1.242</td><td valign="top" align="center" colspan="1">9,2</td></tr><tr><td align="center" colspan="1" valign="top">2</td><td align="left" colspan="1" valign="top"><italic>Mycodiplosis </italic>sp.02</td><td valign="top" align="left" colspan="1">Cecidomyiidae</td><td valign="top" align="center" colspan="1">900</td><td align="center" colspan="1" valign="top">6,7</td></tr><tr><td colspan="1" valign="top" align="center">3</td><td valign="top" align="left" colspan="1"><italic>Culicoides </italic>sp.01</td><td valign="top" align="left" colspan="1">Ceratopogonidae</td><td align="center" colspan="1" valign="top">789</td><td valign="top" align="center" colspan="1">5,8</td></tr><tr><td colspan="1" valign="top" align="center">4</td><td valign="top" align="left" colspan="1"><italic>Miastor </italic>sp.02</td><td valign="top" align="left" colspan="1">Cecidomyiidae</td><td align="center" colspan="1" valign="top">729</td><td align="center" colspan="1" valign="top">5,4</td></tr><tr><td valign="top" align="center" colspan="1">5</td><td valign="top" align="left" colspan="1"><italic>Macrolabis </italic>sp.01</td><td valign="top" align="left" colspan="1">Cecidomyiidae</td><td align="center" colspan="1" valign="top">470</td><td align="center" colspan="1" valign="top">3,5</td></tr><tr><td align="left" colspan="3" valign="top">Kelimpahan lima spesies dominan (<italic>Abundance of five dominant species</italic>)</td><td valign="top" align="center" colspan="1">4.130</td><td valign="top" align="center" colspan="1">30,6</td></tr><tr><td rowspan="6" valign="top" align="left" colspan="1">Hutan alami (<italic>Forest</italic>)</td><td colspan="1" valign="top" align="center">1</td><td valign="top" align="left" colspan="1"><italic>Procontarinia</italic> sp.01</td><td align="left" colspan="1" valign="top">Cecidomyiidae</td><td colspan="1" valign="top" align="center">742</td><td align="center" colspan="1" valign="top">14,5</td></tr><tr><td valign="top" align="center" colspan="1">2</td><td valign="top" align="left" colspan="1"><italic>Mycodiplosis </italic>sp.02</td><td valign="top" align="left" colspan="1">Cecidomyiidae</td><td align="center" colspan="1" valign="top">396</td><td align="center" colspan="1" valign="top">7,7</td></tr><tr><td valign="top" align="center" colspan="1">3</td><td align="left" colspan="1" valign="top"><italic>Asphondylia </italic>sp.03</td><td align="left" colspan="1" valign="top">Cecidomyiidae</td><td colspan="1" valign="top" align="center">294</td><td valign="top" align="center" colspan="1">5,8</td></tr><tr><td valign="top" align="center" colspan="1">4</td><td valign="top" align="left" colspan="1"><italic>Miastor </italic>sp.02</td><td valign="top" align="left" colspan="1">Cecidomyiidae</td><td valign="top" align="center" colspan="1">283</td><td align="center" colspan="1" valign="top">5,5</td></tr><tr><td align="center" colspan="1" valign="top">5</td><td valign="top" align="left" colspan="1"><italic>Culicoides </italic>sp.01</td><td valign="top" align="left" colspan="1">Ceratopogonidae</td><td align="center" colspan="1" valign="top">244</td><td colspan="1" valign="top" align="center">4,8</td></tr><tr><td align="left" colspan="3" valign="top">Kelimpahan lima spesies dominan (<italic>Abundance of five dominant species</italic>)</td><td valign="top" align="center" colspan="1">1.959</td><td valign="top" align="center" colspan="1">38,3</td></tr><tr><td align="left" colspan="1" rowspan="6" valign="top">Hutan karet (<italic>Jungle rubber</italic>)</td><td align="center" colspan="1" valign="top">1</td><td valign="top" align="left" colspan="1"><italic>Procontarinia </italic>sp.01</td><td valign="top" align="left" colspan="1">Cecidomyiidae</td><td valign="top" align="center" colspan="1">390</td><td align="center" colspan="1" valign="top">9,3</td></tr><tr><td valign="top" align="center" colspan="1">2</td><td colspan="1" valign="top" align="left"><italic>Mycodiplosis </italic>sp.02</td><td align="left" colspan="1" valign="top">Cecidomyiidae</td><td valign="top" align="center" colspan="1">379</td><td align="center" colspan="1" valign="top">9,1</td></tr><tr><td valign="top" align="center" colspan="1">3</td><td valign="top" align="left" colspan="1"><italic>Miastor </italic>sp.02</td><td valign="top" align="left" colspan="1">Cecidomyiidae</td><td valign="top" align="center" colspan="1">362</td><td align="center" colspan="1" valign="top">8,7</td></tr><tr><td align="center" colspan="1" valign="top">4</td><td valign="top" align="left" colspan="1"><italic>Culicoides </italic>sp.01</td><td align="left" colspan="1" valign="top">Ceratopogonidae</td><td align="center" colspan="1" valign="top">298</td><td colspan="1" valign="top" align="center">7,1</td></tr><tr><td valign="top" align="center" colspan="1">5</td><td align="left" colspan="1" valign="top"><italic>Macrolabis </italic>sp.01</td><td valign="top" align="left" colspan="1">Cecidomyiidae</td><td colspan="1" valign="top" align="center">270</td><td valign="top" align="center" colspan="1">6,5</td></tr><tr><td valign="top" align="left" colspan="3">Kelimpahan lima spesies dominan (<italic>Abundance of five dominant species</italic>)</td><td valign="top" align="center" colspan="1">1.699</td><td valign="top" align="center" colspan="1">40,7</td></tr><tr><td colspan="1" rowspan="6" valign="top" align="left">Perkebunan karet (<italic>Rubber plantation</italic>)</td><td align="center" colspan="1" valign="top">1</td><td colspan="1" valign="top" align="left"><italic>Brachypogon </italic>sp.01</td><td align="left" colspan="1" valign="top">Ceratopogonidae</td><td valign="top" align="center" colspan="1">216</td><td valign="top" align="center" colspan="1">8,4</td></tr><tr><td align="center" colspan="1" valign="top">2</td><td valign="top" align="left" colspan="1"><italic>Rheotanytarsus </italic>sp.03</td><td valign="top" align="left" colspan="1">Chironomidae</td><td align="center" colspan="1" valign="top">211</td><td align="center" colspan="1" valign="top">8,2</td></tr><tr><td valign="top" align="center" colspan="1">3</td><td valign="top" align="left" colspan="1"><italic>Culicoides </italic>sp.01</td><td align="left" colspan="1" valign="top">Ceratopogonidae</td><td colspan="1" valign="top" align="center">176</td><td align="center" colspan="1" valign="top">6,9</td></tr><tr><td valign="top" align="center" colspan="1">4</td><td colspan="1" valign="top" align="left"><italic>Paramyia </italic>sp.13</td><td align="left" colspan="1" valign="top">Milichiidae</td><td align="center" colspan="1" valign="top">153</td><td align="center" colspan="1" valign="top">6,0</td></tr><tr><td colspan="1" valign="top" align="center">5</td><td colspan="1" valign="top" align="left"><italic>Dasyhelea </italic>sp.01</td><td colspan="1" valign="top" align="left">Ceratopogonidae</td><td align="center" colspan="1" valign="top">111</td><td align="center" colspan="1" valign="top">4,3</td></tr><tr><td valign="top" align="left" colspan="3">Kelimpahan lima spesies dominan (<italic>Abundance of five dominant species</italic>)</td><td colspan="1" valign="top" align="center">867</td><td valign="top" align="center" colspan="1">33,8</td></tr><tr><td colspan="1" rowspan="6" valign="top" align="left">Perkebunan kelapa sawit (<italic>Oil palm plantation</italic>)</td><td valign="top" align="center" colspan="1">1</td><td colspan="1" valign="top" align="left"><italic>Dasyhelea </italic>sp.01</td><td valign="top" align="left" colspan="1">Ceratopogonidae</td><td colspan="1" valign="top" align="center">103</td><td valign="top" align="center" colspan="1">6,2</td></tr><tr><td valign="top" align="center" colspan="1">2</td><td valign="top" align="left" colspan="1"><italic>Culicoides </italic>sp.01</td><td colspan="1" valign="top" align="left">Ceratopogonidae</td><td colspan="1" valign="top" align="center">71</td><td valign="top" align="center" colspan="1">4,3</td></tr><tr><td align="center" colspan="1" valign="top">3</td><td align="left" colspan="1" valign="top"><italic>Gonomyia </italic>sp.01</td><td valign="top" align="left" colspan="1">Limoniidae</td><td align="center" colspan="1" valign="top">71</td><td align="center" colspan="1" valign="top">4,3</td></tr><tr><td align="center" colspan="1" valign="top">4</td><td valign="top" align="left" colspan="1"><italic>Krenosmittia </italic>sp.03</td><td valign="top" align="left" colspan="1">Chironomidae</td><td valign="top" align="center" colspan="1">71</td><td valign="top" align="center" colspan="1">4,3</td></tr><tr><td valign="top" align="center" colspan="1">5</td><td valign="top" align="left" colspan="1"><italic>Forcipomyia </italic>sp.01</td><td valign="top" align="left" colspan="1">Ceratopogonidae</td><td valign="top" align="center" colspan="1">57</td><td valign="top" align="center" colspan="1">3,4</td></tr><tr><td valign="top" align="left" colspan="3">Kelimpahan lima spesies dominan (<italic>Abundance of five dominant species</italic>)</td><td colspan="1" valign="top" align="center">373</td><td align="center" colspan="1" valign="top">22,5</td></tr></tbody></table></table-wrap><table-wrap id="table-3" ignoredToc=""><label>Tabel 3</label><caption><p>Keanekaragaman Diptera di empat tipe penggunaan lahan</p><p>Diversity of Diptera on four land use types</p></caption><table frame="box" rules="all"><thead><tr><th valign="middle" align="center" colspan="1">Tipe penggunaan lahan (<italic>Land use type</italic>)</th><th align="center" colspan="1" valign="middle">Kelimpahan (<italic>Abundance</italic>)</th><th valign="middle" align="center" colspan="1">Famili (<italic>Family</italic>)</th><th align="center" colspan="1" valign="middle">Subfamili (<italic>Subfamily</italic>)</th><th align="center" colspan="1" valign="middle">Genus (<italic>Genera</italic>)</th><th align="center" colspan="1" valign="middle">Spesies (<italic>Species</italic>)</th><th colspan="1" valign="middle" align="center">ACE (<italic>ACE</italic>)</th><th valign="middle" align="center" colspan="1">Spesies dikoleksi (<italic>Collected species</italic>) (%)</th></tr></thead><tbody><tr><td align="left" colspan="1" valign="top">Hutan alami (<italic>Forest</italic>)</td><td align="center" colspan="1" valign="middle">5.113</td><td align="center" colspan="1" valign="middle">34</td><td align="center" colspan="1" valign="middle">59</td><td colspan="1" valign="middle" align="center">129</td><td valign="middle" align="center" colspan="1">287</td><td valign="middle" align="center" colspan="1">369</td><td valign="middle" align="center" colspan="1">77,9</td></tr><tr><td valign="top" align="left" colspan="1">Hutan karet (<italic>Jungle rubber</italic>)</td><td valign="middle" align="center" colspan="1">4.175</td><td align="center" colspan="1" valign="middle">32</td><td align="center" colspan="1" valign="middle">58</td><td valign="middle" align="center" colspan="1">133</td><td align="center" colspan="1" valign="middle">263</td><td valign="middle" align="center" colspan="1">344</td><td valign="middle" align="center" colspan="1">76,5</td></tr><tr><td align="left" colspan="1" valign="top">Perkebunan karet (<italic>Rubber plantation</italic>)</td><td valign="middle" align="center" colspan="1">2.564</td><td align="center" colspan="1" valign="middle">23</td><td valign="middle" align="center" colspan="1">48</td><td valign="middle" align="center" colspan="1">105</td><td valign="middle" align="center" colspan="1">187</td><td valign="middle" align="center" colspan="1">270</td><td valign="middle" align="center" colspan="1">69,2</td></tr><tr><td colspan="1" valign="top" align="left">Perkebunan kelapa sawit (<italic>Oil palm plantation</italic>)</td><td align="center" colspan="1" valign="middle">1.661</td><td align="center" colspan="1" valign="middle">26</td><td align="center" colspan="1" valign="middle">50</td><td align="center" colspan="1" valign="middle">113</td><td colspan="1" valign="middle" align="center">201</td><td valign="middle" align="center" colspan="1">254</td><td colspan="1" valign="middle" align="center">79</td></tr></tbody></table><table-wrap-foot><p>ACE: <italic>abundance-based coverage estimator.</italic></p></table-wrap-foot></table-wrap><fig id="figure-2" ignoredToc=""><label>Gambar 2</label><caption><p>Keterwakilan sampel Diptera di empat tipe penggunaan lahan. A: kurva akumulasi spesies; B: peringkat kelimpahan dari 399 spesies Diptera.</p><p>Representativeness of Diptera samples on four land use types. A: species accumulation curve; B: rank abundance of 399 Diptera species.</p></caption><graphic mime-subtype="jpeg" mimetype="image" xlink:href="https://jurnal.pei-pusat.org/index.php/jei/article/download/890/655/9029"><alt-text>Image</alt-text></graphic></fig><sec><title>Kelimpahan dan kekayaan spesies Diptera di dua lanskap dan empat tipe penggunaan lahan </title><p>Berdasarkan perbedaan lanskap, kelimpahan Diptera lebih tinggi di Hutan Harapan dibandingkan dengan di Bukit Duabelas, yaitu 6.800 dan 6.713 individu. Lebih lanjut, pada masing-masing tipe penggunaan lahan, kelimpahan Diptera kanopi tertinggi ditemukan di lahan hutan alami, yaitu sebanyak 5.113 individu atau 37,8% dari keseluruhan individu yang ditemukan (<xref ref-type="table" rid="table-3">Tabel 3</xref>). Kemudian, kelimpahan tertinggi kedua terdapat pada hutan karet (4.175 individu; 30,9%), diikuti perkebunan karet (2.564 individu; 19%), dan terendah di perkebunan kelapa sawit (1.661 individu; 12,3%) (<xref ref-type="table" rid="table-3">Tabel 3</xref>). Selain memiliki kelimpahan tertinggi, tipe penggunaan lahan hutan alami juga memiliki kekayaan spesies tertinggi, yaitu sebanyak 287 spesies. Kemudian diikuti oleh hutan karet (263 spesies), perkebunan kelapa sawit (201 spesies), dan terendah perkebunan karet (187 spesies) (<xref ref-type="table" rid="table-2">Tabel 2</xref>;<xref ref-type="fig" rid="figure-5">Gambar 3</xref>). Berdasarkan hasil ANOVA, tipe penggunaan lahan berpengaruh nyata terhadap kelimpahan (glm; χ2<sub>(3, 27)</sub> = 23,5; p &lt; 0,001) dan kekayaan spesies Diptera kanopi (glm; F<sub>(3, 27)</sub> = 7,2; p &lt; 0,01), akan tetapi perbedaan lanskap tidak berpengaruh (kelimpahan, glm: χ2(<sub>1,30</sub>) = 0,005; p = 0,9; kekayaan spesies, glm: F(<sub>1,30</sub>) = 0,1; p = 0,7) (<xref ref-type="fig" rid="figure-3">Gambar 4A</xref>, 4B). Hasil ANOVA dari interaksi lanskap dan tipe penggunaan lahan juga menunjukkan hasil yang tidak signifikan terhadap kelimpahan (glm; χ2<sub>(3,24)</sub> = 4,4; p = 0,2) dan kekayaan spesies (glm; F<sub>(3,24)</sub> = 1,5; p = 0,3) (<xref ref-type="fig" rid="figure-3">Gambar 4A</xref>, <xref ref-type="fig" rid="figure-3">4B</xref>). Rata-rata kelimpahan Diptera di hutan alami sekitar dua kali lebih tinggi dibandingkan dengan perkebunan karet dan tiga kali lebih tinggi dibandingkan dengan perkebunan kelapa sawit (<xref ref-type="fig" rid="figure-3">Gambar 5A</xref>). Uji lanjut Tukey menunjukkan bahwa perbedaan tersebut signifikan, sedangkan perbandingan antara hutan alami dan hutan karet tidak menunjukkan perbedaan yang signifikan (<xref ref-type="fig" rid="figure-3">Gambar 5A</xref>). Pola serupa terlihat pada kekayaan spesies Diptera (<xref ref-type="fig" rid="figure-3">Gambar 5B</xref>).</p><fig id="figure-5" ignoredToc=""><label>Gambar 3</label><caption><p>Diptera di empat tipe penggunaan lahan.</p><p>Diptera on four land use types.</p></caption><graphic mime-subtype="png" mimetype="image" xlink:href="https://jurnal.pei-pusat.org/index.php/jei/article/download/890/655/9030"><alt-text>Image</alt-text></graphic></fig><fig ignoredToc="" id="figure-3"><label>Gambar 4</label><caption><p>Pengaruh konversi lahan terhadap Diptera di lanskap Bukit Duabelas dan Hutan Harapan. A: kelimpahan dan B: kekayaan spesies pada empat tipe penggunaan lahan (F: hutan alami; J: hutan karet; R: perkebunan karet; O: perkebunan kelapa sawit). Hasil uji statistik untuk setiap variabel prediktor ditunjukkan sebagai berikut: LS: lanskap, TPL: tipe penggunaan lahan, interaksi: interaksi antara lanskap dan tipe penggunaan lahan, t.s.: tidak signifikan, <italic>: p &lt; 0.01; </italic>*: p &lt; 0.001.</p><p>Effects of land use conversion on Diptera in the Bukit Duabelas and Harapan landscapes. A: abundance and B: species richness across four land use types (F: forest; J: jungle rubber; R: rubber plantation; O: oil palm planta-tion). Results of statistical tests for each predictor variable are presented as follows: LS: landscape; TPL: land-use type; interaction: interaction between landscape and land use type, n.s.: not significant, : p &lt; 0.01, *: p &lt; 0.001.</p></caption><graphic mime-subtype="png" mimetype="image" xlink:href="https://jurnal.pei-pusat.org/index.php/jei/article/download/890/655/9031"><alt-text>Image</alt-text></graphic></fig><fig id="figure-3" ignoredToc=""><label>Gambar 5</label><caption><p>Pengaruh konversi lahan terhadap Diptera. A: kelimpahan dan B: kekayaan spesies (hasil uji Tukey taraf α = 5%). Huruf menunjukkan tipe penggunaan lahan (F: hutan alami; J: hutan karet; R: perkebunan karet; O: perkebunan kelapa sawit).</p><p>Effect of land conversion on Diptera. A: abundance and B: species richness (Tukey test results at α = 5%). The letter indicates the land use type (F: forest; J: jungle rubber; R: rubber plantation; O: oil palm plantation).</p></caption><graphic mime-subtype="png" mimetype="image" xlink:href="https://jurnal.pei-pusat.org/index.php/jei/article/download/890/655/9032"><alt-text>Image</alt-text></graphic></fig></sec><sec><title>Indeks keanekaragam Diptera di empat tipe penggunaan lahan</title><p>Berdasarkan indeks Shannon-Wiener, keempat tipe penggunaan lahan memiliki kompleksitas keanekaragaman Diptera yang tinggi (H’ &gt; 3), yang tertinggi ditemukan pada perkebunan kelapa sawit. Kurva peringkat kelimpahan pada perkebunan kelapa sawit menunjukkan penurunan yang paling landai, menandakan distribusi kelimpahan individu yang lebih merata dan tidak terjadi dominansi spesies tertentu (<xref ref-type="fig" rid="figure-2">Gambar 2</xref>B, <xref ref-type="table" rid="table-4">Tabel 4</xref>). Pola ini sesuai dengan nilai indeks Evenness (E) dan Simpson (1 – D) yang lebih tinggi pada perkebunan kelapa sawit, meskipun kekayaan spesiesnya lebih rendah dibandingkan dengan hutan alami (<xref ref-type="table" rid="table-4">Tabel 4</xref>).</p><table-wrap id="table-4" ignoredToc=""><label>Tabel 4</label><caption><p>Keanekaragaman Diptera di empat tipe penggunaan lahan</p><p>Diversity of Diptera on four land use types</p></caption><table frame="box" rules="all"><thead><tr><th valign="top" align="left" colspan="1">Tipe penggunaan lahan (<italic>Land use type</italic>)</th><th valign="middle" align="center" colspan="1">Spesies (<italic>Species</italic>)</th><th valign="middle" align="center" colspan="1">Shannon-Wiener (H’)</th><th align="center" colspan="1" valign="middle">Simpson (1-D)</th><th colspan="1" valign="middle" align="center">Evenness (E)</th></tr></thead><tbody><tr><td colspan="1" valign="top" align="left">Hutan alami <italic>(Forest)</italic></td><td valign="top" align="center" colspan="1">287</td><td colspan="1" valign="top" align="center">3.59 ab</td><td valign="top" align="center" colspan="1">0.94 ab</td><td align="center" colspan="1" valign="top">0.78 a</td></tr><tr><td align="left" colspan="1" valign="top">Hutan karet <italic>(Jungle rubber)</italic></td><td valign="top" align="center" colspan="1">263</td><td colspan="1" valign="top" align="center">3.56 ab</td><td align="center" colspan="1" valign="top">0.95 ab</td><td valign="top" align="center" colspan="1">0.80 a</td></tr><tr><td colspan="1" valign="top" align="left">Perkebunan karet <italic>(Rubber plantation)</italic></td><td align="center" colspan="1" valign="top">187</td><td valign="top" align="center" colspan="1">3.33 a</td><td align="center" colspan="1" valign="top">0.93 a</td><td valign="top" align="center" colspan="1">0.83 ab</td></tr><tr><td valign="top" align="left" colspan="1">Perkebunan kelapa sawit <italic>(Oil palm plantation)</italic></td><td colspan="1" valign="top" align="center">201</td><td valign="top" align="center" colspan="1">3.71 b</td><td valign="top" align="center" colspan="1">0.96 b</td><td align="center" colspan="1" valign="top">0.89 b</td></tr></tbody></table><table-wrap-foot><p>Huruf yang mengikuti nilai pada setiap baris menunjukkan perbedaan yang signifikan berdasarkan uji Tukey (α = 0,05).</p><p>(Letters following the values in each row indicate significant differences according to Tukey’s test (α = 0.05)).</p></table-wrap-foot></table-wrap></sec><sec><title>Komposisi spesies Diptera di dua lanskap dan empat tipe penggunaan lahan</title><p>Hasil analisis multivariat menunjukkan perbedaan tipe penggunaan lahan (wald<sub>(3,27)</sub> = 44,5; p &lt; 0,001), lanskap (wald<sub>(1,30)</sub> = 23,3; p &lt; 0,001), dan interaksi keduanya (wald<sub>(3,27)</sub> = 25; p &lt; 0,001) memengaruhi komposisi Diptera. Berdasarkan nilai Bray-Curtis, komposisi Diptera pada hutan alami (BF) paling mirip dengan hutan karet (BJ) di lanskap Bukit Duabelas (<xref ref-type="table" rid="table-5">Tabel 5</xref>, <xref ref-type="fig" rid="figure-4">Gambar 6</xref>). Komposisi Diptera yang paling berbeda ditemukan antara hutan alami (BF) dan perkebunan kelapa sawit (BO) di lanskap Bukit Duabelas (<xref ref-type="table" rid="table-5">Tabel 5</xref>, <xref ref-type="fig" rid="figure-4">Gambar 6</xref>). Hasil ini mengindikasikan bahwa <italic>turnover rate</italic> paling rendah terjadi ketika hutan alami beralih menjadi hutan karet. Sebaliknya, <italic>turnover rate</italic> paling tinggi ketika hutan alami dikonversi menjadi perkebunan kelapa sawit. <italic>Turnover</italic> juga dapat dilihat dari pergantian spesies pada setiap tipe penggunaan lahan sebagaimana ditunjukkan <xref ref-type="fig" rid="figure-5">Gambar 3</xref> dan <xref ref-type="table" rid="table-6">Tabel 6</xref>. Lebih lanjut, <xref rid="figure-5" ref-type="fig">Gambar 3</xref> dan <xref ref-type="table" rid="table-7">Tabel 7</xref> menampilkan sebanyak 26% atau 103 spesies yang dijumpai di seluruh tipe penggunaan lahan. Hal ini mengindikasikan adanya spesies generalis yang mampu beradaptasi di berbagai habitat, termasuk habitat yang telah mengalami konversi. Spesies generalis yang paling mendominasi adalah <italic>Procontarinia</italic> sp.01 (Cecidomyiidae) (<xref ref-type="table" rid="table-2">Tabel 2</xref>).</p><table-wrap id="table-5" ignoredToc=""><label>Tabel 5</label><caption><p>Nilai indeks kemiripan Bray-Curtis Diptera di empat tipe penggunaan lahan di lanskap Bukit Duabelas dan Hutan Harapan</p><p>Bray-Curtis similarity index score of Diptera on four land use types in Bukit Duabelas and Harapan Rainforest landscapes</p></caption><table rules="all" frame="box"><thead><tr><th valign="middle" align="center" colspan="1">Tipe penggunaan lahan (<italic>Land use type</italic>)</th><th valign="middle" align="center" colspan="1">BF</th><th align="center" colspan="1" valign="middle">BJ</th><th align="center" colspan="1" valign="middle">BR</th><th valign="middle" align="center" colspan="1">BO</th><th valign="middle" align="center" colspan="1">HF</th><th valign="middle" align="center" colspan="1">HJ</th><th colspan="1" valign="middle" align="center">HR</th><th colspan="1" valign="middle" align="center">HO</th></tr></thead><tbody><tr><td valign="top" align="center" colspan="1">BF</td><td align="center" colspan="1" valign="top">1,00</td><td align="center" colspan="1" valign="top"></td><td colspan="1" valign="top" align="center"></td><td align="center" colspan="1" valign="top"></td><td valign="top" align="center" colspan="1"></td><td valign="top" align="center" colspan="1"></td><td valign="top" align="center" colspan="1"></td><td valign="top" align="center" colspan="1"></td></tr><tr><td valign="top" align="center" colspan="1">BJ</td><td valign="top" align="center" colspan="1">0,60</td><td valign="top" align="center" colspan="1">1,00</td><td valign="top" align="center" colspan="1"></td><td valign="top" align="center" colspan="1"></td><td align="center" colspan="1" valign="top"></td><td align="center" colspan="1" valign="top"></td><td valign="top" align="center" colspan="1"></td><td align="center" colspan="1" valign="top"></td></tr><tr><td align="center" colspan="1" valign="top">BR</td><td align="center" colspan="1" valign="top">0,36</td><td colspan="1" valign="top" align="center">0,41</td><td valign="top" align="center" colspan="1">1,00</td><td align="center" colspan="1" valign="top"></td><td align="center" colspan="1" valign="top"></td><td colspan="1" valign="top" align="center"></td><td colspan="1" valign="top" align="center"></td><td valign="top" align="center" colspan="1"></td></tr><tr><td valign="top" align="center" colspan="1">BO</td><td valign="top" align="center" colspan="1">0,24</td><td valign="top" align="center" colspan="1">0,29</td><td valign="top" align="center" colspan="1">0,44</td><td valign="top" align="center" colspan="1">1,00</td><td valign="top" align="center" colspan="1"></td><td valign="top" align="center" colspan="1"></td><td valign="top" align="center" colspan="1"></td><td align="center" colspan="1" valign="top"></td></tr><tr><td align="center" colspan="1" valign="top">HF</td><td valign="top" align="center" colspan="1">0,53</td><td colspan="1" valign="top" align="center">0,58</td><td colspan="1" valign="top" align="center">0,37</td><td colspan="1" valign="top" align="center">0,30</td><td valign="top" align="center" colspan="1">1,00</td><td valign="top" align="center" colspan="1"></td><td align="center" colspan="1" valign="top"></td><td valign="top" align="center" colspan="1"></td></tr><tr><td valign="top" align="center" colspan="1">HJ</td><td align="center" colspan="1" valign="top">0,52</td><td valign="top" align="center" colspan="1">0,51</td><td valign="top" align="center" colspan="1">0,51</td><td colspan="1" valign="top" align="center">0,41</td><td align="center" colspan="1" valign="top">0,49</td><td align="center" colspan="1" valign="top">1,00</td><td valign="top" align="center" colspan="1"></td><td colspan="1" valign="top" align="center"></td></tr><tr><td align="center" colspan="1" valign="top">HR</td><td valign="top" align="center" colspan="1">0,36</td><td valign="top" align="center" colspan="1">0,33</td><td align="center" colspan="1" valign="top">0,42</td><td align="center" colspan="1" valign="top">0,33</td><td align="center" colspan="1" valign="top">0,27</td><td valign="top" align="center" colspan="1">0,46</td><td valign="top" align="center" colspan="1">1,00</td><td valign="top" align="center" colspan="1"></td></tr><tr><td align="center" colspan="1" valign="top">HO</td><td valign="top" align="center" colspan="1">0,26</td><td valign="top" align="center" colspan="1">0,28</td><td valign="top" align="center" colspan="1">0,43</td><td align="center" colspan="1" valign="top">0,48</td><td align="center" colspan="1" valign="top">0,27</td><td align="center" colspan="1" valign="top">0,43</td><td valign="top" align="center" colspan="1">0,37</td><td colspan="1" valign="top" align="center">1,00</td></tr></tbody></table><table-wrap-foot><p>Huruf pertama menunjukkan lanskap Bukit Duabelas (B) atau Hutan Harapan (H), huruf kedua menunjukkan tipe penggunaan lahan (F: hutan alami; J: hutan karet; R: perkebunan karet; O: perkebunan kelapa sawit).</p><p>(<italic>The first letter indicates the landscape of Bukit Duabelas (B) or Harapan Rainforest (H), the second letter indicates the land use type (F: forest; J: jungle rubber; R: rubber plantation; O: oil palm plantation)).</italic></p></table-wrap-foot></table-wrap><fig id="figure-4" ignoredToc=""><label>Gambar 6</label><caption><p>Komposisi spesies Diptera pada lanskap dan tipe penggunaan lahan yang berbeda. Huruf pertama menunjukkan lanskap Bukit Duabelas (B) atau Hutan Harapan (H), huruf kedua menunjukkan tipe penggunaan lahan (F: hutan alami; J: hutan karet; R: perkebunan karet; O: perkebunan kelapa sawit). Hasil uji statistik mengenai pengaruh varibel prediktor dituliskan sebagai berikut: LS: lanskap; TPL: tipe penggunaan lahan; interaksi: interaksi antara lanskap dan tipe penggunaan lahan, ***: nilai p &lt; 0.001.</p><p>Diptera species composition on different landscapes and land use types. The first letter indicates the landscape of Bukit Duabelas (B) or Harapan Rainforest (H), the second letter indicates the land use type (F: forest; J: jungle rubber;  R:  rubber  plantation;  O:  oil  palm  plantation).  Results  of  statistical  tests  for  each  predictor  variable  are  presented as follows: LS: landscape, TPL: land use type, interaction: interaction between landscape and land use type, *** = p &lt; 0.001.</p></caption><graphic mime-subtype="jpeg" mimetype="image" xlink:href="https://jurnal.pei-pusat.org/index.php/jei/article/download/890/655/9033"><alt-text>Image</alt-text></graphic></fig><table-wrap id="table-6" ignoredToc=""><label>Tabel 6</label><caption><p>Spesies yang dijumpai di tipe penggunaan lahan tertentu</p><p>Species found on certain land use types</p></caption><table frame="box" rules="all"><thead><tr><th align="left" colspan="1" valign="top">Famili (<italic>Family</italic>)</th><th valign="top" align="center" colspan="1">F</th><th align="center" colspan="1" valign="top">J</th><th colspan="1" valign="top" align="center">R</th><th align="center" colspan="1" valign="top">O</th></tr></thead><tbody><tr><td colspan="1" rowspan="2" valign="top" align="left">Asilidae</td><td align="left" colspan="1" valign="top"><italic>Ommatius</italic> sp.01</td><td valign="top" align="left" colspan="1"><italic>Philonicus</italic> sp.01</td><td valign="top" align="left" colspan="1"></td><td align="left" colspan="1" valign="top"></td></tr><tr><td align="left" colspan="1" valign="top"><italic>Philonicus</italic> sp.02</td><td align="left" colspan="1" valign="top"></td><td align="left" colspan="1" valign="top"></td><td valign="top" align="left" colspan="1"></td></tr><tr><td colspan="1" valign="top" align="left">Calliphoridae</td><td valign="top" align="left" colspan="1"></td><td valign="top" align="left" colspan="1"><italic>Rhyncomya flavibasis</italic></td><td valign="top" align="left" colspan="1"></td><td colspan="1" valign="top" align="left"></td></tr><tr><td align="left" colspan="1" valign="top">Cecidomyiidae</td><td valign="top" align="left" colspan="1"></td><td align="left" colspan="1" valign="top"><italic>Didactilomyia</italic> sp.01</td><td colspan="1" valign="top" align="left"></td><td valign="top" align="left" colspan="1"></td></tr><tr><td align="left" colspan="1" valign="top">Celyphidae</td><td align="left" colspan="1" valign="top"></td><td valign="top" align="left" colspan="1"><italic>Acelyphus</italic> sp.01</td><td align="left" colspan="1" valign="top"></td><td valign="top" align="left" colspan="1"></td></tr><tr><td align="left" colspan="1" valign="top">Chamaemyiidae</td><td align="left" colspan="1" valign="top"></td><td valign="top" align="left" colspan="1"><italic>Leucopis</italic> sp.01</td><td colspan="1" valign="top" align="left"></td><td align="left" colspan="1" valign="top"></td></tr><tr><td align="left" colspan="1" valign="top">Ceratopogonidae</td><td valign="top" align="left" colspan="1"><italic>Allohelea </italic>sp.01</td><td align="left" colspan="1" valign="top"></td><td valign="top" align="left" colspan="1"></td><td valign="top" align="left" colspan="1"></td></tr><tr><td valign="top" align="left" colspan="1">Chironomidae</td><td colspan="1" valign="top" align="left"></td><td align="left" colspan="1" valign="top"></td><td align="left" colspan="1" valign="top"></td><td align="left" colspan="1" valign="top"><italic>Polypedilum</italic> sp.02</td></tr><tr><td align="left" colspan="1" rowspan="16" valign="top">Chloropidae</td><td align="left" colspan="1" valign="top"><italic>Chloropidae</italic> sp.01</td><td align="left" colspan="1" valign="top"><italic>Elachiptera</italic> sp.01</td><td align="left" colspan="1" valign="top"><italic>Incertella</italic> sp.02</td><td align="left" colspan="1" valign="top"><italic>Tricimba</italic> sp.02</td></tr><tr><td align="left" colspan="1" valign="top"><italic>Chloropsina</italic> sp.02</td><td align="left" colspan="1" valign="top"><italic>Elachiptera</italic> sp.02</td><td align="left" colspan="1" valign="top"></td><td align="left" colspan="1" valign="top"></td></tr><tr><td align="left" colspan="1" valign="top"><italic>Chloropsina</italic> sp.03</td><td colspan="1" valign="top" align="left"><italic>Gaurax</italic> sp.01</td><td valign="top" align="left" colspan="1"></td><td align="left" colspan="1" valign="top"></td></tr><tr><td valign="top" align="left" colspan="1"><italic>Chloropsina</italic> sp.05</td><td valign="top" align="left" colspan="1"><italic>Incertella</italic> sp.06</td><td align="left" colspan="1" valign="top"></td><td valign="top" align="left" colspan="1"></td></tr><tr><td align="left" colspan="1" valign="top"><italic>Chloropsina</italic> sp.06</td><td colspan="1" valign="top" align="left"><italic>Rhodesiella</italic> sp.02</td><td valign="top" align="left" colspan="1"></td><td align="left" colspan="1" valign="top"></td></tr><tr><td valign="top" align="left" colspan="1"><italic>Chloropsina</italic> sp.07</td><td align="left" colspan="1" valign="top"><italic>Rhodesiella</italic> sp.04</td><td align="left" colspan="1" valign="top"></td><td valign="top" align="left" colspan="1"></td></tr><tr><td colspan="1" valign="top" align="left"><italic>Chloropsina</italic> sp.08</td><td colspan="1" valign="top" align="left"></td><td align="left" colspan="1" valign="top"></td><td valign="top" align="left" colspan="1"></td></tr><tr><td valign="top" align="left" colspan="1"><italic>Dicraeus</italic> sp.03</td><td align="left" colspan="1" valign="top"></td><td align="left" colspan="1" valign="top"></td><td valign="top" align="left" colspan="1"></td></tr><tr><td valign="top" align="left" colspan="1"><italic>Gampsocera</italic> sp.01</td><td valign="top" align="left" colspan="1"></td><td valign="top" align="left" colspan="1"></td><td align="left" colspan="1" valign="top"></td></tr><tr><td valign="top" align="left" colspan="1"><italic>Gaurax</italic> sp.09</td><td valign="top" align="left" colspan="1"></td><td valign="top" align="left" colspan="1"></td><td valign="top" align="left" colspan="1"></td></tr><tr><td align="left" colspan="1" valign="top"><italic>Gaurax</italic> sp.11</td><td align="left" colspan="1" valign="top"></td><td valign="top" align="left" colspan="1"></td><td valign="top" align="left" colspan="1"></td></tr><tr><td valign="top" align="left" colspan="1"><italic>Gaurax</italic> sp.15</td><td align="left" colspan="1" valign="top"></td><td align="left" colspan="1" valign="top"></td><td valign="top" align="left" colspan="1"></td></tr><tr><td align="left" colspan="1" valign="top"><italic>Gaurax</italic> sp.17</td><td align="left" colspan="1" valign="top"></td><td valign="top" align="left" colspan="1"></td><td valign="top" align="left" colspan="1"></td></tr><tr><td valign="top" align="left" colspan="1"><italic>Parectecephala</italic> sp.01</td><td colspan="1" valign="top" align="left"></td><td valign="top" align="left" colspan="1"></td><td colspan="1" valign="top" align="left"></td></tr><tr><td align="left" colspan="1" valign="top"><italic>Rhopalopterum</italic>  sp.06</td><td valign="top" align="left" colspan="1"></td><td valign="top" align="left" colspan="1"></td><td valign="top" align="left" colspan="1"></td></tr><tr><td colspan="1" valign="top" align="left"><italic>Tricimba</italic> sp.01</td><td valign="top" align="left" colspan="1"></td><td valign="top" align="left" colspan="1"></td><td valign="top" align="left" colspan="1"></td></tr><tr><td colspan="1" valign="top" align="left">Culicidae</td><td align="left" colspan="1" valign="top"><italic>Anopheles</italic> sp.03</td><td align="left" colspan="1" valign="top"></td><td valign="top" align="left" colspan="1"></td><td valign="top" align="left" colspan="1"></td></tr><tr><td valign="top" align="left" colspan="1" rowspan="3">Dolichopodidae</td><td align="left" colspan="1" valign="top"><italic>Diaphorus</italic> sp.01</td><td valign="top" align="left" colspan="1"></td><td valign="top" align="left" colspan="1"></td><td valign="top" align="left" colspan="1"><italic>Diaphorus</italic> sp.02</td></tr><tr><td align="left" colspan="1" valign="top"><italic>Sciapus</italic> sp.01</td><td colspan="1" valign="top" align="left"></td><td valign="top" align="left" colspan="1"></td><td align="left" colspan="1" valign="top"><italic>Enlinia</italic> sp.01</td></tr><tr><td align="left" colspan="1" valign="top"><italic>Xanthochlorus</italic> sp.01</td><td valign="top" align="left" colspan="1"></td><td align="left" colspan="1" valign="top"></td><td valign="top" align="left" colspan="1"></td></tr><tr><td colspan="1" rowspan="8" valign="top" align="left">Drosophilidae</td><td align="left" colspan="1" valign="top"><italic>Drosophila</italic> sp.09</td><td align="left" colspan="1" valign="top"><italic>Drosophila</italic> sp.03</td><td valign="top" align="left" colspan="1"></td><td colspan="1" valign="top" align="left"><italic>Leucophenga</italic> sp.04</td></tr><tr><td valign="top" align="left" colspan="1"><italic>Drosophila</italic> sp.11</td><td align="left" colspan="1" valign="top"><italic>Leucophenga</italic> sp.03</td><td align="left" colspan="1" valign="top"></td><td valign="top" align="left" colspan="1"><italic>Mycodrosophila</italic> sp.01</td></tr><tr><td valign="top" align="left" colspan="1"><italic>Drosophila</italic> sp.12</td><td align="left" colspan="1" valign="top"></td><td valign="top" align="left" colspan="1"></td><td align="left" colspan="1" valign="top"></td></tr><tr><td align="left" colspan="1" valign="top"><italic>Drosophila</italic> sp.13</td><td valign="top" align="left" colspan="1"></td><td align="left" colspan="1" valign="top"></td><td align="left" colspan="1" valign="top"></td></tr><tr><td align="left" colspan="1" valign="top"><italic>Drosophila</italic> sp.14</td><td align="left" colspan="1" valign="top"></td><td valign="top" align="left" colspan="1"></td><td valign="top" align="left" colspan="1"></td></tr><tr><td align="left" colspan="1" valign="top"><italic>Drosophilidae</italic> sp.01</td><td valign="top" align="left" colspan="1"></td><td align="left" colspan="1" valign="top"></td><td colspan="1" valign="top" align="left"></td></tr><tr><td align="left" colspan="1" valign="top"><italic>Stegana</italic> sp.01</td><td align="left" colspan="1" valign="top"></td><td valign="top" align="left" colspan="1"></td><td colspan="1" valign="top" align="left"></td></tr><tr><td align="left" colspan="1" valign="top"><italic>Stegana</italic> sp.02</td><td align="left" colspan="1" valign="top"></td><td align="left" colspan="1" valign="top"></td><td valign="top" align="left" colspan="1"></td></tr><tr><td rowspan="3" valign="top" align="left" colspan="1">Empididae</td><td valign="top" align="left" colspan="1"><italic>Empis</italic> sp.01</td><td align="left" colspan="1" valign="top"></td><td align="left" colspan="1" valign="top"></td><td valign="top" align="left" colspan="1"></td></tr><tr><td colspan="1" valign="top" align="left"><italic>Empis</italic> sp.02</td><td valign="top" align="left" colspan="1"></td><td valign="top" align="left" colspan="1"></td><td align="left" colspan="1" valign="top"></td></tr><tr><td valign="top" align="left" colspan="1"><italic>Empis</italic> sp.03</td><td valign="top" align="left" colspan="1"></td><td align="left" colspan="1" valign="top"></td><td valign="top" align="left" colspan="1"></td></tr><tr><td align="left" colspan="1" valign="top">Hybotidae</td><td valign="top" align="left" colspan="1"><italic>Bicellaria</italic> sp.01</td><td align="left" colspan="1" valign="top"></td><td align="left" colspan="1" valign="top"></td><td colspan="1" valign="top" align="left"></td></tr><tr><td align="left" colspan="1" rowspan="2" valign="top">Keroplatidae</td><td valign="top" align="left" colspan="1"></td><td valign="top" align="left" colspan="1"><italic>Macrocera</italic> sp.03</td><td valign="top" align="left" colspan="1"></td><td align="left" colspan="1" valign="top"></td></tr><tr><td valign="top" align="left" colspan="1"></td><td colspan="1" valign="top" align="left"><italic>Macrocera</italic> sp.04</td><td valign="top" align="left" colspan="1"></td><td valign="top" align="left" colspan="1"></td></tr><tr><td align="left" colspan="1" rowspan="4" valign="top">Lauxaniidae</td><td align="left" colspan="1" valign="top"><italic>Homoneura</italic> sp.03</td><td align="left" colspan="1" valign="top"><italic>Homoneura</italic> sp.02</td><td colspan="1" valign="top" align="left"></td><td valign="top" align="left" colspan="1"></td></tr><tr><td align="left" colspan="1" valign="top"><italic>Homoneura</italic> sp.04</td><td valign="top" align="left" colspan="1"><italic>Homoneura</italic> sp.09</td><td valign="top" align="left" colspan="1"></td><td valign="top" align="left" colspan="1"></td></tr><tr><td valign="top" align="left" colspan="1"><italic>Homoneura</italic> sp.06</td><td valign="top" align="left" colspan="1"><italic>Phobeticomyia</italic> sp.01</td><td align="left" colspan="1" valign="top"></td><td align="left" colspan="1" valign="top"></td></tr><tr><td valign="top" align="left" colspan="1"></td><td valign="top" align="left" colspan="1"><italic>Trigonometopus</italic> sp.01</td><td valign="top" align="left" colspan="1"></td><td valign="top" align="left" colspan="1"></td></tr><tr><td colspan="1" rowspan="4" valign="top" align="left">Limoniidae</td><td valign="top" align="left" colspan="1"><italic>Gonempeda</italic> sp.01</td><td valign="top" align="left" colspan="1"><italic>Antocha</italic> sp.02</td><td valign="top" align="left" colspan="1"><italic>Antocha</italic> sp.01</td><td valign="top" align="left" colspan="1"><italic>Antocha</italic> sp.03</td></tr><tr><td valign="top" align="left" colspan="1"></td><td colspan="1" valign="top" align="left"><italic>Dicranomyia</italic> sp.01</td><td align="left" colspan="1" valign="top"><italic>Epiphragma</italic> sp.01</td><td valign="top" align="left" colspan="1"><italic>Libnotes</italic> sp.03</td></tr><tr><td valign="top" align="left" colspan="1"></td><td align="left" colspan="1" valign="top"><italic>Libnotes</italic> sp.02</td><td align="left" colspan="1" valign="top"><italic>Gonomyia</italic> sp.02</td><td colspan="1" valign="top" align="left"><italic>Limonia</italic> sp.01</td></tr><tr><td valign="top" align="left" colspan="1"></td><td valign="top" align="left" colspan="1"><italic>Limoniinae</italic> sp.01</td><td align="left" colspan="1" valign="top"><italic>Limnophilinae</italic> sp.01</td><td valign="top" align="left" colspan="1"></td></tr><tr><td valign="top" align="left" colspan="1">Lonchaeidae</td><td valign="top" align="left" colspan="1"></td><td valign="top" align="left" colspan="1"><italic>Dasiops </italic>sp.03</td><td valign="top" align="left" colspan="1"></td><td colspan="1" valign="top" align="left"><italic>Dasiops </italic>sp.05</td></tr><tr><td valign="top" align="left" colspan="1">Lygistorrhinidae</td><td colspan="1" valign="top" align="left"><italic>Lygistorrhina </italic>sp.01</td><td align="left" colspan="1" valign="top"></td><td valign="top" align="left" colspan="1"></td><td align="left" colspan="1" valign="top"></td></tr><tr><td valign="top" align="left" colspan="1" rowspan="8">Milichiidae</td><td valign="top" align="left" colspan="1"><italic>Neophyllomyza </italic>sp.05</td><td valign="top" align="left" colspan="1"></td><td colspan="1" valign="top" align="left"><italic>Stomosis </italic>sp.02</td><td valign="top" align="left" colspan="1"></td></tr><tr><td align="left" colspan="1" valign="top"><italic>Paramyia </italic>lutea</td><td colspan="1" valign="top" align="left"></td><td valign="top" align="left" colspan="1"></td><td valign="top" align="left" colspan="1"></td></tr><tr><td align="left" colspan="1" valign="top"><italic>Paramyia </italic>sp.02</td><td colspan="1" valign="top" align="left"></td><td align="left" colspan="1" valign="top"></td><td valign="top" align="left" colspan="1"></td></tr><tr><td valign="top" align="left" colspan="1"><italic>Paramyia </italic>sp.06</td><td align="left" colspan="1" valign="top"></td><td valign="top" align="left" colspan="1"></td><td colspan="1" valign="top" align="left"></td></tr><tr><td align="left" colspan="1" valign="top"><italic>Paramyia </italic>sp.14</td><td align="left" colspan="1" valign="top"></td><td align="left" colspan="1" valign="top"></td><td align="left" colspan="1" valign="top"></td></tr><tr><td valign="top" align="left" colspan="1"><italic>Phyllomyza </italic>sp.02</td><td align="left" colspan="1" valign="top"></td><td align="left" colspan="1" valign="top"></td><td colspan="1" valign="top" align="left"></td></tr><tr><td align="left" colspan="1" valign="top"><italic>Phyllomyza </italic>sp.03</td><td valign="top" align="left" colspan="1"></td><td valign="top" align="left" colspan="1"></td><td valign="top" align="left" colspan="1"></td></tr><tr><td colspan="1" valign="top" align="left"><italic>Stomosis </italic>sp.03</td><td valign="top" align="left" colspan="1"></td><td align="left" colspan="1" valign="top"></td><td valign="top" align="left" colspan="1"></td></tr><tr><td colspan="1" valign="top" align="left">Muscidae</td><td align="left" colspan="1" valign="top"><italic>Atherigona </italic>sp.02</td><td align="left" colspan="1" valign="top"><italic>Atherigona </italic>sp.03</td><td align="left" colspan="1" valign="top"><italic>Atherigona </italic>sp.06</td><td align="left" colspan="1" valign="top"><italic>Atherigona </italic>sp.08</td></tr><tr><td colspan="1" valign="top" align="left">Muscidae</td><td valign="top" align="left" colspan="1"><italic>Atherigona </italic>sp.04</td><td valign="top" align="left" colspan="1"><italic>Thricops </italic>sp.02</td><td valign="top" align="left" colspan="1"></td><td valign="top" align="left" colspan="1"></td></tr><tr><td rowspan="2" valign="top" align="left" colspan="1">Mycetophilidae</td><td align="left" colspan="1" valign="top"><italic>Neoempheria </italic>sp.02</td><td colspan="1" valign="top" align="left"><italic>Azana </italic>sp.02</td><td valign="top" align="left" colspan="1"></td><td colspan="1" valign="top" align="left"><italic>Allactoneura </italic>sp.01</td></tr><tr><td valign="top" align="left" colspan="1"></td><td valign="top" align="left" colspan="1">Neoempheria sp.01</td><td align="left" colspan="1" valign="top"></td><td align="left" colspan="1" valign="top"></td></tr><tr><td colspan="1" rowspan="2" valign="top" align="left">Odiniidae</td><td valign="top" align="left" colspan="1"><italic>Odinia </italic>sp.01</td><td valign="top" align="left" colspan="1"></td><td valign="top" align="left" colspan="1"></td><td valign="top" align="left" colspan="1"></td></tr><tr><td align="left" colspan="1" valign="top"><italic>Odinia </italic>sp.02</td><td align="left" colspan="1" valign="top"></td><td colspan="1" valign="top" align="left"></td><td align="left" colspan="1" valign="top"></td></tr><tr><td valign="top" align="left" colspan="1">Pipunculidae</td><td valign="top" align="left" colspan="1"><italic>Pipunculus </italic>sp.02</td><td align="left" colspan="1" valign="top"><italic>Pipunculus </italic>sp.01</td><td valign="top" align="left" colspan="1"></td><td valign="top" align="left" colspan="1"><italic>Tomosvaryella </italic>sp.01</td></tr><tr><td colspan="1" valign="top" align="left">Scatopsidae</td><td align="left" colspan="1" valign="top"><italic>Rhegmoclemina </italic>sp.02</td><td align="left" colspan="1" valign="top"></td><td align="left" colspan="1" valign="top"></td><td colspan="1" valign="top" align="left"></td></tr><tr><td align="left" colspan="1" valign="top">Sciaridae</td><td valign="top" align="left" colspan="1"><italic>Scythropochroa </italic>sp.03</td><td align="left" colspan="1" valign="top"></td><td align="left" colspan="1" valign="top"></td><td colspan="1" valign="top" align="left"><italic>Rhynchosciara </italic>sp.01</td></tr><tr><td colspan="1" valign="top" align="left">Sciomyzidae</td><td align="left" colspan="1" valign="top"></td><td valign="top" align="left" colspan="1"><italic>Sepedomerus </italic>sp.01</td><td valign="top" align="left" colspan="1"></td><td valign="top" align="left" colspan="1"></td></tr><tr><td colspan="1" rowspan="2" valign="top" align="left">Sphaeroceridae</td><td valign="top" align="left" colspan="1"><italic>Poecilosomella </italic>sp.01</td><td valign="top" align="left" colspan="1"></td><td valign="top" align="left" colspan="1"></td><td valign="top" align="left" colspan="1"><italic>Poecilosomella </italic>sp.02</td></tr><tr><td colspan="1" valign="top" align="left"><italic>Terrilimosina </italic>sp.01</td><td align="left" colspan="1" valign="top"></td><td valign="top" align="left" colspan="1"></td><td valign="top" align="left" colspan="1"></td></tr><tr><td colspan="1" rowspan="4" valign="top" align="left">Stratiomyidae</td><td colspan="1" valign="top" align="left"><italic>Gongrosargus </italic>sp.01</td><td valign="top" align="left" colspan="1"><italic>Hermetis </italic>sp.02</td><td align="left" colspan="1" valign="top"><italic>Tinda </italic>sp.02</td><td colspan="1" valign="top" align="left"><italic>Sagaricera </italic>sp.01</td></tr><tr><td colspan="1" valign="top" align="left"><italic>Hermetia </italic>sp.01</td><td align="left" colspan="1" valign="top"><italic>Sargus </italic>sp.01</td><td colspan="1" valign="top" align="left"><italic>Zabrachia </italic>sp.01</td><td align="left" colspan="1" valign="top"></td></tr><tr><td valign="top" align="left" colspan="1"><italic>Isomerocera </italic>sp.01</td><td align="left" colspan="1" valign="top"></td><td align="left" colspan="1" valign="top"></td><td align="left" colspan="1" valign="top"></td></tr><tr><td colspan="1" valign="top" align="left"><italic>Pseudoxymyia </italic>sp.01</td><td colspan="1" valign="top" align="left"></td><td valign="top" align="left" colspan="1"></td><td align="left" colspan="1" valign="top"></td></tr><tr><td colspan="1" rowspan="2" valign="top" align="left">Syrphidae</td><td valign="top" align="left" colspan="1"></td><td align="left" colspan="1" valign="top"><italic>Myolepta </italic>sp.01</td><td valign="top" align="left" colspan="1"></td><td colspan="1" valign="top" align="left"></td></tr><tr><td align="left" colspan="1" valign="top"></td><td valign="top" align="left" colspan="1"><italic>Myolepta </italic>sp.02</td><td valign="top" align="left" colspan="1"></td><td valign="top" align="left" colspan="1"></td></tr><tr><td valign="top" align="left" colspan="1">Tabanidae</td><td align="left" colspan="1" valign="top"><italic>Hamatabanus </italic>sp.01</td><td align="left" colspan="1" valign="top"></td><td colspan="1" valign="top" align="left"></td><td valign="top" align="left" colspan="1"></td></tr><tr><td rowspan="4" valign="top" align="left" colspan="1">Tachinidae</td><td valign="top" align="left" colspan="1"><italic>Tachinidae </italic>sp.02</td><td align="left" colspan="1" valign="top"></td><td valign="top" align="left" colspan="1"></td><td valign="top" align="left" colspan="1"><italic>Tachinidae </italic>sp.01</td></tr><tr><td valign="top" align="left" colspan="1"><italic>Tachinidae </italic>sp.03</td><td colspan="1" valign="top" align="left"></td><td colspan="1" valign="top" align="left"></td><td colspan="1" valign="top" align="left"></td></tr><tr><td valign="top" align="left" colspan="1"><italic>Tachinidae </italic>sp.05</td><td valign="top" align="left" colspan="1"></td><td valign="top" align="left" colspan="1"></td><td align="left" colspan="1" valign="top"></td></tr><tr><td align="left" colspan="1" valign="top"><italic>Tachinidae </italic>sp.06</td><td valign="top" align="left" colspan="1"></td><td align="left" colspan="1" valign="top"></td><td valign="top" align="left" colspan="1"></td></tr><tr><td align="left" colspan="1" valign="top">Tephritidae</td><td align="left" colspan="1" valign="top"></td><td valign="top" align="left" colspan="1"><italic>Dacus </italic>sp.01</td><td valign="top" align="left" colspan="1"><italic>Anastrepha </italic>sp.01</td><td valign="top" align="left" colspan="1"></td></tr><tr><td valign="top" align="left" colspan="1">Tephritidae</td><td valign="top" align="left" colspan="1"></td><td valign="top" align="left" colspan="1"></td><td valign="top" align="left" colspan="1"><italic>Cornutrypeta </italic>sp.01</td><td valign="top" align="left" colspan="1"></td></tr><tr><td align="left" colspan="1" valign="top">Xylophagidae</td><td valign="top" align="left" colspan="1"><italic>Dialysis </italic>sp.01</td><td align="left" colspan="1" valign="top"></td><td colspan="1" valign="top" align="left"></td><td colspan="1" valign="top" align="left"></td></tr></tbody></table><table-wrap-foot><p>F: hutan alami <italic>(forest)</italic>; J: hutan karet <italic>(jungle rubber)</italic>; R: perkebunan karet <italic>(rubber plantation)</italic>; O: perkebunan kelapa sawit <italic>(oil palm plantation).</italic></p></table-wrap-foot></table-wrap><table-wrap id="table-7" ignoredToc=""><label>Tabel 7</label><caption><p>Sebanyak 103 spesies generalis dari 18 famili di empat tipe penggunaan lahan </p><p>A total of 103 generalist species from 18 families on four land use types</p></caption><table rules="all" frame="box"><thead><tr><th valign="top" align="left" colspan="1">Famili (<italic>Family</italic>)</th><th valign="top" align="center" colspan="1">Spesies (<italic>Species</italic>)</th><th align="center" colspan="1" valign="top">F</th><th align="center" colspan="1" valign="top">J</th><th align="center" colspan="1" valign="top">R</th><th valign="top" align="center" colspan="1">O</th></tr></thead><tbody><tr><td rowspan="22" valign="top" align="left" colspan="1">Cecidomyiidae</td><td align="left" colspan="1" valign="top"><italic>Asphondylia </italic>sp.01</td><td valign="top" align="center" colspan="1">23</td><td colspan="1" valign="top" align="center">52</td><td valign="top" align="center" colspan="1">4</td><td valign="top" align="center" colspan="1">7</td></tr><tr><td align="left" colspan="1" valign="top"><italic>Asphondylia </italic>sp.03</td><td valign="top" align="center" colspan="1">294</td><td valign="top" align="center" colspan="1">23</td><td align="center" colspan="1" valign="top">10</td><td colspan="1" valign="top" align="center">7</td></tr><tr><td colspan="1" valign="top" align="left"><italic>Camptomyia </italic>sp.01</td><td valign="top" align="center" colspan="1">55</td><td valign="top" align="center" colspan="1">33</td><td valign="top" align="center" colspan="1">7</td><td valign="top" align="center" colspan="1">4</td></tr><tr><td valign="top" align="left" colspan="1"><italic>Didactilomyia </italic>sp.02</td><td colspan="1" valign="top" align="center">87</td><td valign="top" align="center" colspan="1">13</td><td valign="top" align="center" colspan="1">12</td><td valign="top" align="center" colspan="1">17</td></tr><tr><td valign="top" align="left" colspan="1"><italic>Feltiella </italic>sp.01</td><td valign="top" align="center" colspan="1">12</td><td align="center" colspan="1" valign="top">4</td><td align="center" colspan="1" valign="top">8</td><td valign="top" align="center" colspan="1">5</td></tr><tr><td align="left" colspan="1" valign="top"><italic>Feltiella </italic>sp.02</td><td align="center" colspan="1" valign="top">14</td><td align="center" colspan="1" valign="top">24</td><td valign="top" align="center" colspan="1">11</td><td align="center" colspan="1" valign="top">3</td></tr><tr><td valign="top" align="left" colspan="1"><italic>Giardomyia </italic>sp.01</td><td colspan="1" valign="top" align="center">2</td><td align="center" colspan="1" valign="top">7</td><td align="center" colspan="1" valign="top">5</td><td align="center" colspan="1" valign="top">1</td></tr><tr><td align="left" colspan="1" valign="top"><italic>Heteropeza </italic>sp.01</td><td valign="top" align="center" colspan="1">2</td><td valign="top" align="center" colspan="1">2</td><td valign="top" align="center" colspan="1">1</td><td valign="top" align="center" colspan="1">3</td></tr><tr><td align="left" colspan="1" valign="top"><italic>Lestremia </italic>sp.01</td><td colspan="1" valign="top" align="center">4</td><td colspan="1" valign="top" align="center">9</td><td align="center" colspan="1" valign="top">3</td><td valign="top" align="center" colspan="1">2</td></tr><tr><td valign="top" align="left" colspan="1"><italic>Macrolabis </italic>sp.01</td><td valign="top" align="center" colspan="1">136</td><td colspan="1" valign="top" align="center">270</td><td valign="top" align="center" colspan="1">50</td><td valign="top" align="center" colspan="1">14</td></tr><tr><td valign="top" align="left" colspan="1"><italic>Miastor </italic>sp.01</td><td colspan="1" valign="top" align="center">5</td><td valign="top" align="center" colspan="1">5</td><td align="center" colspan="1" valign="top">6</td><td valign="top" align="center" colspan="1">12</td></tr><tr><td align="left" colspan="1" valign="top"><italic>Miastor </italic>sp.02</td><td align="center" colspan="1" valign="top">283</td><td align="center" colspan="1" valign="top">362</td><td align="center" colspan="1" valign="top">44</td><td colspan="1" valign="top" align="center">40</td></tr><tr><td valign="top" align="left" colspan="1"><italic>Micromya </italic>sp.01</td><td align="center" colspan="1" valign="top">21</td><td align="center" colspan="1" valign="top">31</td><td valign="top" align="center" colspan="1">20</td><td valign="top" align="center" colspan="1">42</td></tr><tr><td valign="top" align="left" colspan="1"><italic>Monardia </italic>sp.01</td><td valign="top" align="center" colspan="1">4</td><td valign="top" align="center" colspan="1">17</td><td valign="top" align="center" colspan="1">2</td><td valign="top" align="center" colspan="1">6</td></tr><tr><td align="left" colspan="1" valign="top"><italic>Mycodiplosis </italic>sp.01</td><td valign="top" align="center" colspan="1">16</td><td valign="top" align="center" colspan="1">31</td><td colspan="1" valign="top" align="center">45</td><td align="center" colspan="1" valign="top">9</td></tr><tr><td align="left" colspan="1" valign="top"><italic>Mycodiplosis </italic>sp.02</td><td valign="top" align="center" colspan="1">396</td><td align="center" colspan="1" valign="top">379</td><td align="center" colspan="1" valign="top">79</td><td valign="top" align="center" colspan="1">46</td></tr><tr><td valign="top" align="left" colspan="1"><italic>Peromyia </italic>sp.01</td><td valign="top" align="center" colspan="1">48</td><td align="center" colspan="1" valign="top">70</td><td valign="top" align="center" colspan="1">15</td><td valign="top" align="center" colspan="1">21</td></tr><tr><td align="left" colspan="1" valign="top"><italic>Procontarinia </italic>sp.01</td><td valign="top" align="center" colspan="1">742</td><td valign="top" align="center" colspan="1">390</td><td valign="top" align="center" colspan="1">67</td><td align="center" colspan="1" valign="top">43</td></tr><tr><td valign="top" align="left" colspan="1"><italic>Rhopalomyia </italic>sp.01</td><td align="center" colspan="1" valign="top">38</td><td valign="top" align="center" colspan="1">14</td><td colspan="1" valign="top" align="center">12</td><td align="center" colspan="1" valign="top">3</td></tr><tr><td align="left" colspan="1" valign="top"><italic>Vitisiella </italic>sp.01</td><td valign="top" align="center" colspan="1">94</td><td align="center" colspan="1" valign="top">24</td><td align="center" colspan="1" valign="top">7</td><td align="center" colspan="1" valign="top">7</td></tr><tr><td colspan="1" valign="top" align="left"><italic>Winnertzia </italic>sp.01</td><td colspan="1" valign="top" align="center">25</td><td align="center" colspan="1" valign="top">41</td><td align="center" colspan="1" valign="top">10</td><td align="center" colspan="1" valign="top">9</td></tr><tr><td valign="top" align="left" colspan="1"><italic>Winnertzia </italic>sp.02</td><td align="center" colspan="1" valign="top">9</td><td valign="top" align="center" colspan="1">6</td><td align="center" colspan="1" valign="top">9</td><td align="center" colspan="1" valign="top">6</td></tr><tr><td rowspan="19" valign="top" align="left" colspan="1">Ceratopogonidae</td><td align="left" colspan="1" valign="top"><italic>Allohelea johannseni</italic></td><td valign="top" align="center" colspan="1">23</td><td align="center" colspan="1" valign="top">74</td><td valign="top" align="center" colspan="1">3</td><td colspan="1" valign="top" align="center">2</td></tr><tr><td valign="top" align="left" colspan="1"><italic>Alluaudomyia </italic>sp.01</td><td align="center" colspan="1" valign="top">10</td><td align="center" colspan="1" valign="top">11</td><td colspan="1" valign="top" align="center">1</td><td valign="top" align="center" colspan="1">2</td></tr><tr><td align="left" colspan="1" valign="top"><italic>Alluaudomyia </italic>sp.02</td><td valign="top" align="center" colspan="1">7</td><td valign="top" align="center" colspan="1">23</td><td align="center" colspan="1" valign="top">16</td><td valign="top" align="center" colspan="1">4</td></tr><tr><td valign="top" align="left" colspan="1"><italic>Alluaudomyia </italic>sp.03</td><td align="center" colspan="1" valign="top">9</td><td valign="top" align="center" colspan="1">41</td><td colspan="1" valign="top" align="center">33</td><td align="center" colspan="1" valign="top">4</td></tr><tr><td align="left" colspan="1" valign="top"><italic>Alluaudomyia </italic>sp.04</td><td align="center" colspan="1" valign="top">2</td><td align="center" colspan="1" valign="top">2</td><td align="center" colspan="1" valign="top">12</td><td valign="top" align="center" colspan="1">5</td></tr><tr><td valign="top" align="left" colspan="1"><italic>Atrichopogon </italic>sp.01</td><td align="center" colspan="1" valign="top">21</td><td align="center" colspan="1" valign="top">23</td><td valign="top" align="center" colspan="1">15</td><td valign="top" align="center" colspan="1">31</td></tr><tr><td align="left" colspan="1" valign="top"><italic>Bezzia </italic>sp.01</td><td valign="top" align="center" colspan="1">4</td><td align="center" colspan="1" valign="top">4</td><td valign="top" align="center" colspan="1">1</td><td align="center" colspan="1" valign="top">3</td></tr><tr><td colspan="1" valign="top" align="left"><italic>Bezzia </italic>sp.02</td><td valign="top" align="center" colspan="1">5</td><td align="center" colspan="1" valign="top">5</td><td align="center" colspan="1" valign="top">1</td><td colspan="1" valign="top" align="center">1</td></tr><tr><td colspan="1" valign="top" align="left"><italic>Brachypogon </italic>sp.01</td><td colspan="1" valign="top" align="center">81</td><td valign="top" align="center" colspan="1">120</td><td colspan="1" valign="top" align="center">216</td><td align="center" colspan="1" valign="top">43</td></tr><tr><td align="left" colspan="1" valign="top"><italic>Culicoides </italic>sp.01</td><td valign="top" align="center" colspan="1">244</td><td valign="top" align="center" colspan="1">298</td><td align="center" colspan="1" valign="top">176</td><td valign="top" align="center" colspan="1">71</td></tr><tr><td colspan="1" valign="top" align="left"><italic>Culicoides </italic>sp.02</td><td valign="top" align="center" colspan="1">62</td><td valign="top" align="center" colspan="1">22</td><td valign="top" align="center" colspan="1">25</td><td valign="top" align="center" colspan="1">4</td></tr><tr><td colspan="1" valign="top" align="left"><italic>Culicoides </italic>sp.03</td><td valign="top" align="center" colspan="1">34</td><td valign="top" align="center" colspan="1">10</td><td align="center" colspan="1" valign="top">3</td><td valign="top" align="center" colspan="1">5</td></tr><tr><td colspan="1" valign="top" align="left"><italic>Culicoides </italic>sp.04</td><td align="center" colspan="1" valign="top">154</td><td colspan="1" valign="top" align="center">81</td><td align="center" colspan="1" valign="top">9</td><td colspan="1" valign="top" align="center">26</td></tr><tr><td valign="top" align="left" colspan="1"><italic>Dasyhelea </italic>sp.01</td><td valign="top" align="center" colspan="1">64</td><td valign="top" align="center" colspan="1">103</td><td align="center" colspan="1" valign="top">111</td><td align="center" colspan="1" valign="top">103</td></tr><tr><td valign="top" align="left" colspan="1"><italic>Forcipomyia </italic>sp.01</td><td align="center" colspan="1" valign="top">46</td><td valign="top" align="center" colspan="1">51</td><td align="center" colspan="1" valign="top">31</td><td align="center" colspan="1" valign="top">57</td></tr><tr><td valign="top" align="left" colspan="1"><italic>Heteromyia </italic>sp.01</td><td align="center" colspan="1" valign="top">2</td><td align="center" colspan="1" valign="top">1</td><td valign="top" align="center" colspan="1">2</td><td align="center" colspan="1" valign="top">1</td></tr><tr><td align="left" colspan="1" valign="top"><italic>Monohelea maculipennis</italic></td><td align="center" colspan="1" valign="top">96</td><td valign="top" align="center" colspan="1">27</td><td valign="top" align="center" colspan="1">64</td><td valign="top" align="center" colspan="1">1</td></tr><tr><td valign="top" align="left" colspan="1"><italic>Stenoxenus </italic>sp.01</td><td align="center" colspan="1" valign="top">4</td><td valign="top" align="center" colspan="1">6</td><td valign="top" align="center" colspan="1">18</td><td align="center" colspan="1" valign="top">5</td></tr><tr><td valign="top" align="left" colspan="1"><italic>Stilobezzia </italic>sp.03</td><td align="center" colspan="1" valign="top">114</td><td align="center" colspan="1" valign="top">116</td><td valign="top" align="center" colspan="1">91</td><td align="center" colspan="1" valign="top">25</td></tr><tr><td align="left" colspan="1" rowspan="11" valign="top">Chironomidae</td><td align="left" colspan="1" valign="top"><italic>Corynoneura </italic>sp.01</td><td align="center" colspan="1" valign="top">2</td><td valign="top" align="center" colspan="1">15</td><td valign="top" align="center" colspan="1">19</td><td valign="top" align="center" colspan="1">4</td></tr><tr><td valign="top" align="left" colspan="1"><italic>Derotanypus </italic>sp.01</td><td valign="top" align="center" colspan="1">3</td><td align="center" colspan="1" valign="top">2</td><td valign="top" align="center" colspan="1">1</td><td align="center" colspan="1" valign="top">4</td></tr><tr><td valign="top" align="left" colspan="1"><italic>Glyptotendipes </italic>sp.02</td><td valign="top" align="center" colspan="1">8</td><td colspan="1" valign="top" align="center">14</td><td valign="top" align="center" colspan="1">27</td><td align="center" colspan="1" valign="top">6</td></tr><tr><td align="left" colspan="1" valign="top"><italic>Glyptotendipes </italic>sp.03</td><td valign="top" align="center" colspan="1">14</td><td valign="top" align="center" colspan="1">12</td><td valign="top" align="center" colspan="1">68</td><td align="center" colspan="1" valign="top">11</td></tr><tr><td valign="top" align="left" colspan="1"><italic>Glyptotendipes </italic>sp.04</td><td valign="top" align="center" colspan="1">2</td><td valign="top" align="center" colspan="1">3</td><td align="center" colspan="1" valign="top">23</td><td align="center" colspan="1" valign="top">2</td></tr><tr><td align="left" colspan="1" valign="top"><italic>Krenosmittia </italic>sp.01</td><td colspan="1" valign="top" align="center">3</td><td colspan="1" valign="top" align="center">1</td><td valign="top" align="center" colspan="1">2</td><td valign="top" align="center" colspan="1">4</td></tr><tr><td valign="top" align="left" colspan="1"><italic>Krenosmittia </italic>sp.02</td><td valign="top" align="center" colspan="1">5</td><td valign="top" align="center" colspan="1">2</td><td colspan="1" valign="top" align="center">18</td><td align="center" colspan="1" valign="top">20</td></tr><tr><td valign="top" align="left" colspan="1"><italic>Lasiodiamesa </italic>sp.01</td><td valign="top" align="center" colspan="1">7</td><td valign="top" align="center" colspan="1">25</td><td align="center" colspan="1" valign="top">4</td><td align="center" colspan="1" valign="top">3</td></tr><tr><td valign="top" align="left" colspan="1"><italic>Lasiodiamesa </italic>sp.02</td><td valign="top" align="center" colspan="1">9</td><td align="center" colspan="1" valign="top">18</td><td valign="top" align="center" colspan="1">8</td><td align="center" colspan="1" valign="top">6</td></tr><tr><td valign="top" align="left" colspan="1"><italic>Lasiodiamesa </italic>sp.03</td><td colspan="1" valign="top" align="center">58</td><td valign="top" align="center" colspan="1">22</td><td align="center" colspan="1" valign="top">20</td><td align="center" colspan="1" valign="top">4</td></tr><tr><td valign="top" align="left" colspan="1"><italic>Lasiodiamesa </italic>sp.04</td><td align="center" colspan="1" valign="top">2</td><td valign="top" align="center" colspan="1">1</td><td valign="top" align="center" colspan="1">1</td><td align="center" colspan="1" valign="top">2</td></tr><tr><td align="left" colspan="1" rowspan="5" valign="top"></td><td valign="top" align="left" colspan="1"><italic>Polypedilum </italic>sp.01</td><td align="center" colspan="1" valign="top">32</td><td valign="top" align="center" colspan="1">41</td><td valign="top" align="center" colspan="1">64</td><td align="center" colspan="1" valign="top">4</td></tr><tr><td align="left" colspan="1" valign="top"><italic>Rheotanytarsus </italic>sp.01</td><td valign="top" align="center" colspan="1">13</td><td valign="top" align="center" colspan="1">38</td><td align="center" colspan="1" valign="top">80</td><td valign="top" align="center" colspan="1">19</td></tr><tr><td valign="top" align="left" colspan="1"><italic>Rheotanytarsus </italic>sp.02</td><td valign="top" align="center" colspan="1">2</td><td valign="top" align="center" colspan="1">7</td><td align="center" colspan="1" valign="top">57</td><td valign="top" align="center" colspan="1">8</td></tr><tr><td valign="top" align="left" colspan="1"><italic>Rheotanytarsus </italic>sp.03</td><td align="center" colspan="1" valign="top">22</td><td valign="top" align="center" colspan="1">44</td><td valign="top" align="center" colspan="1">211</td><td valign="top" align="center" colspan="1">10</td></tr><tr><td align="left" colspan="1" valign="top"><italic>Rheotanytarsus </italic>sp.04</td><td valign="top" align="center" colspan="1">1</td><td align="center" colspan="1" valign="top">5</td><td align="center" colspan="1" valign="top">2</td><td align="center" colspan="1" valign="top">1</td></tr><tr><td align="left" colspan="1" rowspan="7" valign="top"></td><td valign="top" align="left" colspan="1"><italic>Symbiocladius </italic>sp.01</td><td colspan="1" valign="top" align="center">7</td><td colspan="1" valign="top" align="center">8</td><td valign="top" align="center" colspan="1">4</td><td valign="top" align="center" colspan="1">5</td></tr><tr><td align="left" colspan="1" valign="top"><italic>Symbiocladius </italic>sp.02</td><td colspan="1" valign="top" align="center">1</td><td align="center" colspan="1" valign="top">1</td><td valign="top" align="center" colspan="1">9</td><td valign="top" align="center" colspan="1">1</td></tr><tr><td valign="top" align="left" colspan="1"><italic>Thienemanniella </italic>sp.01</td><td align="center" colspan="1" valign="top">24</td><td valign="top" align="center" colspan="1">6</td><td colspan="1" valign="top" align="center">8</td><td align="center" colspan="1" valign="top">5</td></tr><tr><td valign="top" align="left" colspan="1"><italic>Thienemanniella </italic>sp.02</td><td valign="top" align="center" colspan="1">2</td><td colspan="1" valign="top" align="center">4</td><td align="center" colspan="1" valign="top">27</td><td valign="top" align="center" colspan="1">47</td></tr><tr><td valign="top" align="left" colspan="1"><italic>Thienemanniella </italic>sp.03</td><td valign="top" align="center" colspan="1">100</td><td valign="top" align="center" colspan="1">55</td><td align="center" colspan="1" valign="top">17</td><td valign="top" align="center" colspan="1">13</td></tr><tr><td align="left" colspan="1" valign="top"><italic>Zavrelia </italic>sp.01</td><td valign="top" align="center" colspan="1">19</td><td align="center" colspan="1" valign="top">15</td><td valign="top" align="center" colspan="1">25</td><td valign="top" align="center" colspan="1">4</td></tr><tr><td align="left" colspan="1" valign="top"><italic>Zavrelia </italic>sp.02</td><td align="center" colspan="1" valign="top">60</td><td align="center" colspan="1" valign="top">24</td><td valign="top" align="center" colspan="1">21</td><td colspan="1" valign="top" align="center">4</td></tr><tr><td valign="top" align="left" colspan="1" rowspan="3">Chloropidae</td><td colspan="1" valign="top" align="left"><italic>Gaurax </italic>sp.12</td><td align="center" colspan="1" valign="top">61</td><td align="center" colspan="1" valign="top">17</td><td valign="top" align="center" colspan="1">4</td><td align="center" colspan="1" valign="top">1</td></tr><tr><td valign="top" align="left" colspan="1"><italic>Gaurax </italic>sp.14</td><td valign="top" align="center" colspan="1">3</td><td align="center" colspan="1" valign="top">12</td><td valign="top" align="center" colspan="1">6</td><td valign="top" align="center" colspan="1">4</td></tr><tr><td valign="top" align="left" colspan="1"><italic>Gaurax </italic>sp.16</td><td align="center" colspan="1" valign="top">7</td><td align="center" colspan="1" valign="top">4</td><td valign="top" align="center" colspan="1">9</td><td valign="top" align="center" colspan="1">2</td></tr><tr><td valign="top" align="left" colspan="1">Culicidae</td><td valign="top" align="left" colspan="1"><italic>Mimomyia </italic>sp.02</td><td align="center" colspan="1" valign="top">9</td><td align="center" colspan="1" valign="top">4</td><td align="center" colspan="1" valign="top">3</td><td align="center" colspan="1" valign="top">1</td></tr><tr><td align="left" colspan="1" rowspan="6" valign="top">Dolichopodidae</td><td valign="top" align="left" colspan="1"><italic>Chrysotus </italic>sp.01</td><td align="center" colspan="1" valign="top">1</td><td colspan="1" valign="top" align="center">5</td><td colspan="1" valign="top" align="center">2</td><td valign="top" align="center" colspan="1">3</td></tr><tr><td colspan="1" valign="top" align="left"><italic>Diaphorus </italic>sp.03</td><td valign="top" align="center" colspan="1">9</td><td valign="top" align="center" colspan="1">11</td><td colspan="1" valign="top" align="center">9</td><td colspan="1" valign="top" align="center">10</td></tr><tr><td valign="top" align="left" colspan="1"><italic>Sciapus </italic>sp.02</td><td valign="top" align="center" colspan="1">10</td><td colspan="1" valign="top" align="center">7</td><td align="center" colspan="1" valign="top">9</td><td align="center" colspan="1" valign="top">17</td></tr><tr><td valign="top" align="left" colspan="1"><italic>Sciapus </italic>sp.04</td><td align="center" colspan="1" valign="top">7</td><td valign="top" align="center" colspan="1">12</td><td align="center" colspan="1" valign="top">5</td><td colspan="1" valign="top" align="center">13</td></tr><tr><td valign="top" align="left" colspan="1"><italic>Thrypticus </italic>sp.01</td><td valign="top" align="center" colspan="1">2</td><td align="center" colspan="1" valign="top">13</td><td valign="top" align="center" colspan="1">16</td><td colspan="1" valign="top" align="center">6</td></tr><tr><td valign="top" align="left" colspan="1"><italic>Xanthochlorus</italic> sp.05</td><td align="center" colspan="1" valign="top">1</td><td align="center" colspan="1" valign="top">2</td><td valign="top" align="center" colspan="1">2</td><td colspan="1" valign="top" align="center">1</td></tr><tr><td align="left" colspan="1" rowspan="2" valign="top">Drosophilidae</td><td valign="top" align="left" colspan="1"><italic>Drosophila </italic>sp.04</td><td align="center" colspan="1" valign="top">3</td><td align="center" colspan="1" valign="top">3</td><td align="center" colspan="1" valign="top">1</td><td valign="top" align="center" colspan="1">7</td></tr><tr><td valign="top" align="left" colspan="1"><italic>Drosophila </italic>sp.10</td><td valign="top" align="center" colspan="1">2</td><td colspan="1" valign="top" align="center">14</td><td valign="top" align="center" colspan="1">18</td><td valign="top" align="center" colspan="1">29</td></tr><tr><td align="left" colspan="1" valign="top">Ephydridae</td><td align="left" colspan="1" valign="top"><italic>Ptilomyia </italic>sp.01</td><td valign="top" align="center" colspan="1">25</td><td align="center" colspan="1" valign="top">5</td><td align="center" colspan="1" valign="top">1</td><td valign="top" align="center" colspan="1">3</td></tr><tr><td align="left" colspan="1" valign="top">Keroplatidae</td><td valign="top" align="left" colspan="1"><italic>Macrocera </italic>sp.01</td><td valign="top" align="center" colspan="1">2</td><td align="center" colspan="1" valign="top">2</td><td valign="top" align="center" colspan="1">5</td><td valign="top" align="center" colspan="1">3</td></tr><tr><td align="left" colspan="1" valign="top">Lauxaniidae</td><td valign="top" align="left" colspan="1"><italic>Sapromyza </italic>sp.01</td><td align="center" colspan="1" valign="top">6</td><td valign="top" align="center" colspan="1">2</td><td colspan="1" valign="top" align="center">1</td><td align="center" colspan="1" valign="top">2</td></tr><tr><td valign="top" align="left" colspan="1" rowspan="2">Limoniidae</td><td valign="top" align="left" colspan="1"><italic>Gonomyia </italic>sp.01</td><td align="center" colspan="1" valign="top">4</td><td valign="top" align="center" colspan="1">8</td><td valign="top" align="center" colspan="1">14</td><td colspan="1" valign="top" align="center">71</td></tr><tr><td align="left" colspan="1" valign="top"><italic>Trentepohlia </italic>(<italic>Monogoma</italic>) sp.01</td><td colspan="1" valign="top" align="center">2</td><td valign="top" align="center" colspan="1">16</td><td valign="top" align="center" colspan="1">5</td><td align="center" colspan="1" valign="top">19</td></tr><tr><td align="left" colspan="1" valign="top">Lonchaeidae</td><td valign="top" align="left" colspan="1"><italic>Dasiops </italic>sp.04</td><td colspan="1" valign="top" align="center">7</td><td valign="top" align="center" colspan="1">5</td><td valign="top" align="center" colspan="1">1</td><td valign="top" align="center" colspan="1">1</td></tr><tr><td align="left" colspan="1" rowspan="5" valign="top">Milichiidae</td><td align="left" colspan="1" valign="top"><italic>Neophyllomyza </italic>sp.02</td><td align="center" colspan="1" valign="top">7</td><td align="center" colspan="1" valign="top">4</td><td valign="top" align="center" colspan="1">2</td><td align="center" colspan="1" valign="top">7</td></tr><tr><td align="left" colspan="1" valign="top"><italic>Paramyia </italic>sp.04</td><td colspan="1" valign="top" align="center">4</td><td align="center" colspan="1" valign="top">8</td><td valign="top" align="center" colspan="1">5</td><td valign="top" align="center" colspan="1">17</td></tr><tr><td align="left" colspan="1" valign="top"><italic>Paramyia </italic>sp.09</td><td valign="top" align="center" colspan="1">10</td><td align="center" colspan="1" valign="top">3</td><td colspan="1" valign="top" align="center">1</td><td valign="top" align="center" colspan="1">11</td></tr><tr><td align="left" colspan="1" valign="top"><italic>Paramyia </italic>sp.10</td><td colspan="1" valign="top" align="center">29</td><td valign="top" align="center" colspan="1">13</td><td valign="top" align="center" colspan="1">46</td><td align="center" colspan="1" valign="top">16</td></tr><tr><td valign="top" align="left" colspan="1"><italic>Paramyia </italic>sp.13</td><td valign="top" align="center" colspan="1">108</td><td valign="top" align="center" colspan="1">64</td><td valign="top" align="center" colspan="1">153</td><td align="center" colspan="1" valign="top">15</td></tr><tr><td align="left" colspan="1" rowspan="5" valign="top">Phoridae</td><td valign="top" align="left" colspan="1"><italic>Dohrniphora </italic>sp.01</td><td valign="top" align="center" colspan="1">3</td><td valign="top" align="center" colspan="1">14</td><td align="center" colspan="1" valign="top">1</td><td valign="top" align="center" colspan="1">3</td></tr><tr><td align="left" colspan="1" valign="top"><italic>Metopininae </italic>sp.01</td><td align="center" colspan="1" valign="top">6</td><td valign="top" align="center" colspan="1">33</td><td valign="top" align="center" colspan="1">12</td><td valign="top" align="center" colspan="1">39</td></tr><tr><td colspan="1" valign="top" align="left"><italic>Metopininae </italic>sp.02</td><td valign="top" align="center" colspan="1">26</td><td valign="top" align="center" colspan="1">12</td><td align="center" colspan="1" valign="top">30</td><td colspan="1" valign="top" align="center">23</td></tr><tr><td valign="top" align="left" colspan="1"><italic>Metopininae </italic>sp.04</td><td align="center" colspan="1" valign="top">2</td><td align="center" colspan="1" valign="top">9</td><td align="center" colspan="1" valign="top">1</td><td align="center" colspan="1" valign="top">24</td></tr><tr><td align="left" colspan="1" valign="top"><italic>Metopininae </italic>sp.05</td><td align="center" colspan="1" valign="top">3</td><td align="center" colspan="1" valign="top">23</td><td colspan="1" valign="top" align="center">15</td><td valign="top" align="center" colspan="1">28</td></tr><tr><td colspan="1" rowspan="2" valign="top" align="left">Psychodidae</td><td align="left" colspan="1" valign="top"><italic>Lutzomyia </italic>sp.01</td><td align="center" colspan="1" valign="top">49</td><td align="center" colspan="1" valign="top">24</td><td align="center" colspan="1" valign="top">2</td><td align="center" colspan="1" valign="top">1</td></tr><tr><td valign="top" align="left" colspan="1"><italic>Telmatoscopus </italic>sp.01</td><td valign="top" align="center" colspan="1">26</td><td valign="top" align="center" colspan="1">11</td><td valign="top" align="center" colspan="1">1</td><td align="center" colspan="1" valign="top">3</td></tr><tr><td rowspan="2" valign="top" align="left" colspan="1">Scatopsidae</td><td align="left" colspan="1" valign="top"><italic>Colobostema </italic>sp.01</td><td valign="top" align="center" colspan="1">1</td><td align="center" colspan="1" valign="top">1</td><td valign="top" align="center" colspan="1">3</td><td colspan="1" valign="top" align="center">2</td></tr><tr><td colspan="1" valign="top" align="left"><italic>Holoplagia </italic>sp.01</td><td align="center" colspan="1" valign="top">5</td><td valign="top" align="center" colspan="1">3</td><td align="center" colspan="1" valign="top">3</td><td align="center" colspan="1" valign="top">2</td></tr><tr><td align="left" colspan="1" rowspan="6" valign="top">Sciaridae</td><td valign="top" align="left" colspan="1"><italic>Bradysia </italic>sp.01</td><td align="center" colspan="1" valign="top">64</td><td align="center" colspan="1" valign="top">44</td><td align="center" colspan="1" valign="top">14</td><td valign="top" align="center" colspan="1">39</td></tr><tr><td align="left" colspan="1" valign="top"><italic>Scatopsciara </italic>sp.03</td><td valign="top" align="center" colspan="1">201</td><td colspan="1" valign="top" align="center">54</td><td align="center" colspan="1" valign="top">8</td><td valign="top" align="center" colspan="1">28</td></tr><tr><td align="left" colspan="1" valign="top"><italic>Scatopsciara </italic>sp.05</td><td valign="top" align="center" colspan="1">15</td><td valign="top" align="center" colspan="1">10</td><td valign="top" align="center" colspan="1">2</td><td colspan="1" valign="top" align="center">4</td></tr><tr><td valign="top" align="left" colspan="1"><italic>Scatopsciara </italic>sp.06</td><td align="center" colspan="1" valign="top">13</td><td align="center" colspan="1" valign="top">8</td><td valign="top" align="center" colspan="1">1</td><td valign="top" align="center" colspan="1">2</td></tr><tr><td valign="top" align="left" colspan="1"><italic>Scatopsciara </italic>sp.07</td><td align="center" colspan="1" valign="top">19</td><td valign="top" align="center" colspan="1">11</td><td align="center" colspan="1" valign="top">2</td><td valign="top" align="center" colspan="1">4</td></tr><tr><td valign="top" align="left" colspan="1"><italic>Scythropochroa </italic>sp.01</td><td align="center" colspan="1" valign="top">5</td><td align="center" colspan="1" valign="top">4</td><td valign="top" align="center" colspan="1">10</td><td align="center" colspan="1" valign="top">7</td></tr><tr><td colspan="1" valign="top" align="left">Tephritidae</td><td align="left" colspan="1" valign="top"><italic>Urophora </italic>sp.01</td><td colspan="1" valign="top" align="center">1</td><td align="center" colspan="1" valign="top">3</td><td align="center" colspan="1" valign="top">1</td><td valign="top" align="center" colspan="1">5</td></tr></tbody></table><table-wrap-foot><p>F: hutan alami <italic>(forest)</italic>; J: hutan karet <italic>(jungle rubber)</italic>; R: perkebunan karet <italic>(rubber plantation)</italic>; O: perkebunan kelapa sawit <italic>(oil palm plantation).</italic></p></table-wrap-foot></table-wrap></sec></sec><sec><title>PEMBAHASAN</title><p>Konversi hutan menjadi kawasan perkebunan monokultur, khususnya karet dan kelapa sawit berdampak signifikan terhadap keberadaan Diptera pada suatu ekosistem. Penelitian ini menunjukkan bahwa kawasan hutan alami dan agroforestri hutan karet dapat mendukung kelimpahan dan kekayaan spesies Diptera yang lebih tinggi dibandingkan dengan kawasan perkebunan karet dan perkebunan kelapa sawit. Pola ini mengindikasikan Diptera merespons perubahan kompleksitas vegetasi yang menjadi habitatnya. Konversi hutan menjadi perkebunan monokultur berarti melakukan penyederhanaan struktur vegetasi. Pengaruh vegetasi menjadi pembahasan penting mengingat objek yang diteliti adalah Diptera yang berasosiasi dengan kanopi pohon.</p><p>Kawasan hutan alami dan hutan karet relatif mirip, yaitu memiliki kompleksitas vegetasi yang tinggi, lapisan kanopi yang rapat, dan intensitas pengolahan lahan yang rendah bahkan tidak sama sekali<xref ref-type="bibr" rid="BIBR-18">(Drescher et al., 2016)</xref>;<xref ref-type="bibr" rid="BIBR-59">(Rembold et al., 2017)</xref>. Vegetasi di hutan alami paling kompleks, dengan kekayaan spesies 1,5 kali lebih tinggi dibandingkan dengan hutan karet dan 4 kali lebih tinggi dibandingkan dengan perkebunan karet dan perkebunan kelapa sawit<xref ref-type="bibr" rid="BIBR-59">(Rembold et al., 2017)</xref>. Kompleksitas vegetasi ini berhubungan positif dengan kekayaan spesies artropoda<xref ref-type="bibr" rid="BIBR-7">(Basset et al., 2012)</xref>. Habitat yang kompleks berarti menyediakan banyak ceruk (<italic>niche</italic>) dengan sumber daya berupa pakan yang beragam, juga tempat berlindung dan melakukan oviposisi<xref ref-type="bibr" rid="BIBR-54">(Prince et al., 2011)</xref>. Keanekaragaman ceruk mendukung kelimpahan dan kekayaan spesies Diptera yang lebih tinggi di hutan dibandingkan dengan perkebunan, sejalan dengan penelitian Geeraert et al. (2018)<xref ref-type="bibr" rid="BIBR-24">(Geeraert et al., 2018)</xref>. Kondisi ini juga memungkinan keberadaan spesies Diptera dengan ceruk yang spesifik dan terspesialisasi sehingga hanya ditemukan di hutan alami dan hutan karet. Berdasarkan laporan Rembold et al. (2017)<xref ref-type="bibr" rid="BIBR-59">(Rembold et al., 2017)</xref>, terdapat 13 spesies tumbuhan yang hanya ditemukan di hutan alami dan hutan karet. Salah satu spesies tumbuhan tersebut kemungkinan merupakan inang dari <italic>Asphondylia</italic> sp.02, yaitu spesies dari Cecidomyiidae yang hanya ditemukan di kedua tipe penggunaan lahan ini. Maia (2024)<xref ref-type="bibr" rid="BIBR-40">(Maia, 2024)</xref> melaporkan, setiap spesies dari Genus <italic>Asphondylia</italic> menginduksi puru pada tumbuhan inang yang spesifik, yaitu <italic>Asphondylia braziliensis</italic> sp. nov. pada <italic>Struthanthus acuminatus</italic> (Ruiz. &amp; Pav.) Kuijt (Loranthaceae), <italic>Asphondylia fluminensis</italic> sp. nov. pada <italic>Erythroxylum ovalifolium</italic> Peyr (Erythroxylaceae), <italic>Asphondylia marambaiensis</italic> sp. nov. pada <italic>Lantana fucata</italic> Lindl. (Verbenaceae), <italic>Asphondylia varroniae</italic> sp. nov. pada <italic>Varronia curassavica</italic> Jacq. (Cordiaceae), dan <italic>Asphondylia xerezi</italic> sp. nov. pada <italic>Heliotropium</italic> sp. (Heliotropiaceae).</p><p>Tutupan kanopi di hutan alami paling rapat dibandingkan dengan tiga tipe penggunaan lahan lainnya sehingga suhu udara lebih rendah dan kelembapan lebih tinggi<xref ref-type="bibr" rid="BIBR-18">(Drescher et al., 2016)</xref>. Lingkungan yang lembap dan intensitas cahaya rendah meningkatkan kelimpahan dan kekayaan spesies sebagian famili dari Diptera<xref ref-type="bibr" rid="BIBR-67">(Toledo-Hernandez et al., 2021)</xref>. Lingkungan yang ternaungi dan tidak terekspos sinar matahari secara langsung, yaitu di dalam kanopi pohon menjadi lokasi <italic>lek</italic> bagi kelompok Diptera jantan<xref ref-type="bibr" rid="BIBR-35">(Kaspi &amp; Yuval, 1999)</xref>;<xref ref-type="bibr" rid="BIBR-21">(Field et al., 2003)</xref>. <italic>Lek</italic> bertujuan untuk menarik serangga betina yang siap kawin<xref ref-type="bibr" rid="BIBR-54">(Prince et al., 2011)</xref>. Famili Diptera yang melakukan <italic>lek</italic> antara lain Drosophilidae, Tephritidae, Platystomatidae, Ephydridae, Micropezidae, Tabanidae, Therevidae, dan Bombyliidae. (Marshall 2017)</p><p>Kerapatan tutupan kanopi memengaruhi keanekaragaman tumbuhan di hutan, yang secara tidak langsung memengaruhi keberadaan Diptera. Namun demikian, keberadaan suatu spesies tumbuhan tidak berdampak seragam terhadap seluruh Diptera, mengingat ceruk ekologi Diptera sangat beragam dan tidak semua spesies Diptera adalah fitofag (Marshall et al. 2017). Scherber et al. (2014)<xref ref-type="bibr" rid="BIBR-60">(Scherber et al., 2014)</xref> melaporkan, tutupan kanopi yang rapat dapat membatasi keberadaan tumbuhan herba sehingga menurunkan kelimpahan dan kekayaan Superfamili Empidoidea dan Famili Phoridae. Keberadaan herba lebih kuat pengaruhnya terhadap Empidoidea karena menjadi tempat Hybotidae predator mencari mangsa dan sebagian anggota Empididae merupakan serangga pengunjung herba berbunga<xref ref-type="bibr" rid="BIBR-60">(Scherber et al., 2014)</xref>. Namun demikian, pola yang dilaporkan Scherber et al. (2014)<xref ref-type="bibr" rid="BIBR-60">(Scherber et al., 2014)</xref> tersebut berbeda dengan hasil penelitian yang kami lakukan. Menurut Scherber et al. (2014)<xref ref-type="bibr" rid="BIBR-60">(Scherber et al., 2014)</xref>, penentuan strata saat pengambilan sampel berpengaruh terhadap hasil, kelimpahan, dan kekayaan spesies Diptera yang diperoleh dari strata tumbuhan herba di atas permukaan tanah akan berbeda dengan yang diperoleh dari kanopi pohon. Perbedaan hasil tersebut karena persebaran spesies Diptera di hutan bervariasi pada setiap strata, dengan proporsi 49% di permukaan tanah, 64% di kanopi pohon, dan 27% di atas kanopi pohon<xref ref-type="bibr" rid="BIBR-16">(Vertical stratification of insect abundance and species richness in an Amazonian tropical forest, 2022)</xref>. Perbedaan persebaran spesies Diptera setiap strata tersebut diduga memengaruhi keterwakilan sampel dalam penelitian kami yang hanya mencapai 87,4% dari metode pengasapan kanopi pohon. Penelitian selanjutnya perlu mempertimbangkan penambahan jumlah plot dan menerapkan pengambilan sampel multistrata.</p><p>Tutupan kanopiyang rapat di hutan alamimembatasi keberadaan tumbuhan epifit yang menempel pada pohon sehingga kelimpahannya rendah<xref ref-type="bibr" rid="BIBR-13">(Böhnert et al., 2016)</xref>. Kelimpahan epifit yang paling tinggi ditemukan di perkebunan kelapa sawit dengan kanopi terbuka<xref ref-type="bibr" rid="BIBR-13">(Böhnert et al., 2016)</xref>. Keberadaan tumbuhan epifit di perkebunan kelapa sawit mendukung kelimpahan dan kekayaan spesies serangga kanopi karena menyediakan habitat dan sumber makanan<xref rid="BIBR-64" ref-type="bibr">(Suzanti et al., 2016)</xref>. Epifit merupakan habitat bagi larva Ceratopogonidae karena kondisinya lembap dan terdapat badan air di dalamnya<xref ref-type="bibr" rid="BIBR-37">(Kirk-Spriggs et al., 2017)</xref>. Sejalan dengan temuan kami di perkebunan kelapa sawit, Ceratopogonidae menunjukkan kelimpahan paling tinggi dibandingkan dengan 25 famili lainnya. <italic>Dasyhelea</italic> sp.01 adalah spesies yang paling dominan di perkebunan kelapa sawit, kemungkinan karena bromelia epifit adalah salah satu <italic>breeding site</italic> yang baik bagi <italic>Dasyhelea</italic><xref ref-type="bibr" rid="BIBR-70">(Winder, 1977)</xref>.</p><p>Kelimpahan Ceratopogonidae yang tinggi di perkebunan karet tidak berkaitan dengan keberadaan epifit. Epifit tidak dapat berkembang dengan baik di pohon karet karena struktur batangnya relatif halus, berbeda dengan batang kelapa sawit yang tersusun atas pelepah-pelepah yang mengakumulasi bahan organik dan mendukung keberadaan epifit<xref ref-type="bibr" rid="BIBR-13">(Böhnert et al., 2016)</xref>. Kelimpahan Ceratopogonidae diduga berhubungan dengan keberadaan pohon karet. Sesuai laporan <xref ref-type="bibr" rid="BIBR-37">(Kirk-Spriggs et al., 2017)</xref>, beberapa spesies dari Ceratopogonidae merupakan polinator penting bagi tanaman karet. Hasil penelitian kami menunjukkan bahwa kelimpahan Ceratopogonidae di perkebunan karet dua kali lebih tinggi dibandingkan dengan perkebunan kelapa sawit. Selain itu, pada sesama lahan hutan, kelimpahan Ceratopogonidae lebih tinggi di hutan karet dibandingkan dengan hutan alami. Ceratopogonidae yang paling dominan di perkebunan karet adalah <italic>Brachypogon</italic> sp.01. Namun demikian, <italic>Brachypogon</italic> bukan merupakan polinator tumbuhan karet, melainkan predator dari Chironomidae<xref ref-type="bibr" rid="BIBR-37">(Kirk-Spriggs et al., 2017)</xref>. Tingginya kelimpahan <italic>Brachypogon</italic> sp.01 kemungkinan berhubungan dengan ketersediaan mangsa, yaitu Chironomidae yang memang paling melimpah di perkebunan karet dibandingkan dengan tipe penggunaan lahan lainnya.</p><p>Dominasi Ceratopogonidae di kawasan perkebunan mengindikasikan terjadinya peralihan famili dominan jika hutan dikonversi menjadi perkebunan. Di kawasan hutan, Cecidomyiidae merupakan famili yang paling dominan, lalu dominasi beralih ke Ceratopogonidae di kawasan perkebunan. Namun demikian, pada tingkat ordo, pola umum tetap menunjukkan penurunan kelimpahan dan kekayaan spesies Diptera di perkebunan, sebagaimana terjadi pada serangga lain seperti semut, kupu-kupu, tawon parasitoid, dan kumbang<xref ref-type="bibr" rid="BIBR-47">(Nazarreta et al., 2020)</xref>.</p><p>Penurunan kelimpahan dan kekayaan spesies Diptera di kawasan perkebunan berkaitan dengan intensifikasi lahan. Aktivitas pengelolaan lahan meliputi pengaplikasian pupuk dan pestisida kimia, pengambilan getah karet, dan pemanenan tandan kelapa sawit<xref ref-type="bibr" rid="BIBR-1">(Allen et al., 2015)</xref>. Aktivitas tersebut dapat mengganggu populasi serangga sehingga menurunkan kelimpahan dan kekayaan spesies<xref ref-type="bibr" rid="BIBR-28">(Haneda &amp; Yuniar, 2015)</xref>. Penurunan kelimpahan dan kekayaan spesies semakin meningkat seiring bertambahnya intensitas dan frekuensi pengelolaan lahan<xref ref-type="bibr" rid="BIBR-28">(Haneda &amp; Yuniar, 2015)</xref>.</p><p>Kelimpahan Diptera di perkebunan kelapa sawit paling rendah, namun memiliki kompleksitas keanekaragaman yang paling tinggi karena tidak ada spesies yang mendominasi secara ekstrem. Kelimpahan setiap spesies Diptera di kawasan ini cenderung merata. Spesies Diptera dengan kelimpahan paling tinggi, yaitu <italic>Dasyhelea</italic> sp.01 (Ceratopogonidae) yang hanya mencakup 6% dari keseluruhan Diptera di kawasan ini. Aktivitas pemanenan dan pemangkasan pelepah secara berkala menyebabkan perubahan struktur kanopi yang bersifat temporer<xref ref-type="bibr" rid="BIBR-27">(Grass et al., 2020)</xref>. Kondisi ini diduga menyebabkan mikroklimat yang dinamis di kanopi kelapa sawit sehingga menghambat terbentuknya dominansi spesies Diptera di kawasan ini. Berbeda dengan perkebunan, intervensi manusia di kawasan hutan alami relatif terbatas sehingga kondisi mikroklimat lebih stabil dan dominansi spesies tertentu lebih tinggi, yaitu <italic>Procontarinia</italic> sp.01 (Cecidomyiidae) dengan kelimpahan mencapai 15%.</p><p><italic>Procontarinia</italic> sp.01 ditemukan di semua tipe penggunaan lahan, yang berarti spesies tersebut bersifat generalis dan mampu menyebar dengan baik. <italic>Procontarinia</italic> merupakan Diptera fitofag penyebab puru pada daun dan bunga yang memiliki rentang distribusi yang luas dan plastisitas ekologi sehingga dapat ditemukan pada berbagai tipe penggunaan lahan sepanjang tahun<xref rid="BIBR-2" ref-type="bibr">(Amouroux et al., 2013)</xref>;<xref ref-type="bibr" rid="BIBR-63">(Susila et al., 2022)</xref>. Plastisitas ekologi yang baik merupakan ciri khas dari spesies Diptera generalis sehingga mendukung kemampuannya untuk beradaptasi bahkan mendominasi di berbagai habitat, serta membedakannya dari spesies spesialis<xref rid="BIBR-8" ref-type="bibr">(Boudreau et al., 2021)</xref>.</p><p>Konversi lahan mendorong terjadinya perubahan komposisi Diptera. Berdasarkan indeks Bray-Curtis, perubahan komposisi paling kecil ditemukan ketika hutan alami dikonversi menjadi hutan karet, yaitu sebesar 40% sebagaimana yang terjadi pada kumbang daun<xref ref-type="bibr" rid="BIBR-3">(Amrulloh et al., 2022)</xref>. Struktur vegetasi hutan alami dan hutan karet tidak jauh berbeda sehingga komposisi Diptera cenderung mirip. Sebaliknya, perubahan komposisi yang paling besar terjadi ketika hutan alami dikonversi menjadi perkebunan kelapa sawit, dengan tingkat perubahan mencapai 76%. Perubahan komposisi Diptera ini didorong oleh Spesies <italic>turnover</italic>. <italic>Turnover rate</italic> pada perkebunan kelapa sawit lebih tinggi, kemungkinan karena struktur vegetasi lebih homogen dan tingkat intensifikasi lahan lebih besar dibandingkan dengan hutan karet, sejalan dengan laporan <xref ref-type="bibr" rid="BIBR-22">(M.Á et al., 2015)</xref>.</p><p>Konversi menyebabkan sebanyak 135 spesies Diptera dari hutan alami hilang dan digantikan oleh 49 spesies pendatang di perkebunan kelapa sawit. Spesies Diptera yang hilang didominasi oleh Famili Chloropidae, Drosophilidae, dan Milichiidae, berturut-turut sebanyak 37, 13, dan 12 spesies. Sebaliknya, spesies pendatang terutama termasuk dalam dari Famili Limoniidae, Dolichopodidae, Chironomidae, dan Milichiidae, masing-masing sebanyak 8, 6, 4, dan 4 spesies. Pada famili lain juga dijumpai spesies yang hilang dan spesies pendatang, namun jumlah spesies tersebut relatif sedikit. Spesies <italic>turnover</italic> ini dapat terjadi karena setiap spesies memiliki sensitivitas, kemampuan adaptasi, dan toleransi yang bervariasi terhadap dinamika kondisi lingkungan<xref ref-type="bibr" rid="BIBR-65">(Taradipha et al., 2018)</xref>. Spesies Diptera yang hilang di perkebunan kelapa sawit kemungkinan adalah spesies spesialis dari hutan alami yang memiliki ceruk spesifik dan ruang penyebaran terbatas sehingga tidak ditemukan di kawasan ini, sejalan dengan laporan Daawia &amp; Dianingsih (2023)<xref ref-type="bibr" rid="BIBR-15">(Daawia &amp; Dianingsih, 2023)</xref>. Spesies Diptera pendatang kemungkinan adalah spesies yang toleran terhadap tekanan lingkungan, yaitu aktivitas pengelolaan lahan secara intensif di perkebunan kelapa sawit yang mencakup homogenisasi vegetasi, pemanenan, pemangkasan, pemupukan, dan penyemprotan insektisida<xref ref-type="bibr" rid="BIBR-44">(Mohd-Azlan et al., 2023)</xref>.</p><p>Diptera merupakan konsumen primer dan konsumen sekunder dalam jejaring makanan, sedangkan tumbuhan berperan sebagai produsen<xref ref-type="bibr" rid="BIBR-36">(Kirk-Spriggs et al., 2017)</xref>. Interaksi antarkomponen dalam jejaring makanan biasanya sudah membentuk hubungan yang erat sehingga hilangnya satu komponen akan menimbulkan efek domino, yaitu turut hilangnya komponen pada tingkat trofik berikutnya<xref ref-type="bibr" rid="BIBR-61">(Schowalter, 2011)</xref>. Jonsson et al. (2006)<xref ref-type="bibr" rid="BIBR-32">(Jonsson et al., 2006)</xref> menguraikan bahwa hilangnya konsumen primer pada struktur jejaring makanan yang sederhana lebih berisiko terhadap kepunahan konsumen sekunder, yang pada akhirnya mengancam kestabilan ekosistem. Jika konversi hutan menjadi kawasan perkebunan terus berlanjut, diperkirakan ekosistem menjadi semakin homogen dan lebih rentan terhadap gangguan<xref ref-type="bibr" rid="BIBR-68">(Laliberté &amp; Tylianakis, 2010)</xref>. Upaya konservasi yang dapat dilakukan salah satunya melalui pengayaan pada kawasan perkebunan, yaitu menanam berbagai jenis tanaman lokal di di dalamnya, sehingga dapat meningkatkan keanekaragaman hayati dan mempertahankan fungsi ekologi<xref rid="BIBR-66" ref-type="bibr">(Teuscher et al., 2016)</xref>;<xref ref-type="bibr" rid="BIBR-6">(Azhar et al., 2022)</xref>;<xref ref-type="bibr" rid="BIBR-71">(Zemp et al., 2019)</xref>;<xref ref-type="bibr" rid="BIBR-72">(Zemp et al., 2023)</xref>, sekaligus meningkatkan hasil panen di lahan tersebut<xref ref-type="bibr" rid="BIBR-25">(Gérard et al., 2017)</xref>.</p></sec><sec><title>KESIMPULAN</title><p>Konversi hutan menjadi perkebunan monokultur menyebabkan penurunan kelimpahan dan kekayaan spesies Diptera kanopi secara signifikan. Kelimpahan terendah ditemukan di perkebunan kelapa sawit, sedangkan kekayaan spesies terendah ditemukan di perkebunan karet. Konversi lahan juga menyebabkan peralihan dominansi, kawasan hutan didominasi oleh Cecidomyiidae, lalu dominasi beralih ke Ceratopogonidae di kawasan perkebunan monokultur. Perubahan komposisi Diptera yang paling besar terjadi ketika hutan alami dikonversi menjadi perkebunan kelapa sawit. Perubahan komposisi ini didorong oleh spesies <italic>turnover,</italic> yaitu hilangnya spesies tertentu dan digantikan oleh spesies pendatang. Namun demikian, terdapat spesies yang tetap ditemukan di seluruh tipe penggunaan dengan kelimpahan tinggi, yaitu <italic>Procontarinia</italic> sp.01. Penelitian ini menyoroti pentingnya keanekaragaman tumbuhan dan kompleksitas struktur kanopi bagi komunitas Diptera. Upaya konservasi melalui pengayaan tumbuhan di kawasan perkebunan monokultur disarankan sebagai langkah untuk meningkatkan keanekaragaman hayati dan mempertahankan fungsi ekologi. Langkah ini tidak hanya penting untuk mempertahankan keanekaragaman Diptera, tetapi juga spesies tumbuhan dan binatang lain yang saling terkait dalam jejaring makanan di ekosistem.</p></sec></body><back><ack><sec><title>UCAPAN TERIMA KASIH</title><p>Terima kasih diucapkan kepada <italic>Deutsche Forschungsgemeinschaft Germany </italic>(DFG) <italic>Research Foundation </italic>yang telah memberikan dukungan pendanaan CRC990/EFForTS ( <italic>Ecological and Socioeconomic Function of Tropical Lowland Rainforest Transformation Systems </italic>) atas kerja sama penelitian yang diberikan. Terima kasih diucapkan kepada Lembaga Pengelola Dana Pendidikan (LPDP) yang telah memberikan dukungan pendanaan studi.</p></sec></ack><ref-list><title>REFERENCES</title><ref id="BIBR-1"><element-citation publication-type="article-journal"><article-title>Soil nitrogen-cycling responses to conversion of lowland forests to oil palm and rubber plantations in Sumatra, Indonesia</article-title><source>Public Library of Science One</source><volume>10</volume><person-group person-group-type="author"><name><surname>Allen</surname><given-names>K.</given-names></name><name><surname>Corre</surname><given-names>M.D.</given-names></name><name><surname>Tjoa</surname><given-names>A.</given-names></name><name><surname>Veldkamp</surname><given-names>E.</given-names></name></person-group><year>2015</year><fpage>1</fpage><lpage>21</lpage><page-range>1-21</page-range><pub-id pub-id-type="doi">10.1371/journal.pone.0133325</pub-id></element-citation></ref><ref id="BIBR-2"><element-citation publication-type="article-journal"><article-title>Invasive mango blossom gall midge, Procontarinia mangiferae (Felt) (Diptera: Cecidomyiidae) in Reunion Island: ecological plasticity, permanent and structured populations</article-title><source>Biological Invasions</source><volume>15</volume><person-group person-group-type="author"><name><surname>Amouroux</surname><given-names>P.</given-names></name><name><surname>Normand</surname><given-names>F.</given-names></name><name><surname>Nibouche</surname><given-names>S.</given-names></name><name><surname>Delatte</surname><given-names>H.</given-names></name></person-group><year>2013</year><fpage>1677</fpage><lpage>1693</lpage><page-range>1677-1693</page-range><pub-id pub-id-type="doi">10.1007/s10530-012-0400-0</pub-id></element-citation></ref><ref id="BIBR-3"><element-citation publication-type="article-journal"><article-title>Keanekaragaman dan kelimpahan kumbang daun (Coleoptera: Chrysomelidae) pada empat tipe penggunaan lahan yang berbeda di Taman Nasional Bukit Duabelas dan Hutan Harapan, Provinsi Jambi</article-title><source>Jurnal Entomologi Indonesia</source><volume>19</volume><person-group person-group-type="author"><name><surname>Amrulloh</surname><given-names>R.</given-names></name><name><surname>Noerdjito</surname><given-names>W.A.</given-names></name><name><surname>Istiaji</surname><given-names>B.</given-names></name><name><surname>Hidayat</surname><given-names>P.</given-names></name><name><surname>Buchori</surname><given-names>D.</given-names></name></person-group><year>2022</year><fpage>147</fpage><lpage>163</lpage><page-range>147-163</page-range><pub-id pub-id-type="doi">10.5994/jei.19.2.147</pub-id></element-citation></ref><ref id="BIBR-4"><element-citation publication-type="article-journal"><article-title>Rating knockdown of flour beetles after exposure to two insecticides as an indicator of mortality</article-title><source>Scientific Reports</source><volume>11</volume><person-group person-group-type="author"><name><surname>Athanassiou</surname><given-names>C.G.</given-names></name><name><surname>Kavallieratos</surname><given-names>N.G.</given-names></name><name><surname>Arthur</surname><given-names>F.H.</given-names></name><name><surname>Nakas</surname><given-names>C.T.</given-names></name></person-group><year>2021</year><pub-id pub-id-type="doi">10.1038/s41598-020-78982-z</pub-id></element-citation></ref><ref id="BIBR-5"><element-citation publication-type="article-journal"><article-title>Rainforest conversion to cash crops reduces abundance, biomass and species richness of parasitoid wasps in Sumatra, Indonesia</article-title><source>Agricultural and Forest Entomology</source><volume>24</volume><person-group person-group-type="author"><name><surname>Azhar</surname><given-names>A.</given-names></name><name><surname>Hartke</surname><given-names>T.R.</given-names></name><name><surname>Böttges</surname><given-names>L.</given-names></name><name><surname>Lang</surname><given-names>T.</given-names></name><name><surname>Larasati</surname><given-names>A.</given-names></name><name><surname>Novianti</surname><given-names>N.</given-names></name><name><surname>Tawakkal</surname><given-names>I.</given-names></name><name><surname>Hidayat</surname><given-names>P.</given-names></name><name><surname>Buchori</surname><given-names>D.</given-names></name><name><surname>Scheu</surname><given-names>S.</given-names></name><name><surname>Drescher</surname><given-names>J.</given-names></name></person-group><year>2022</year><fpage>506</fpage><lpage>515</lpage><page-range>506-515</page-range><pub-id pub-id-type="doi">10.1111/afe.12512</pub-id></element-citation></ref><ref id="BIBR-6"><element-citation publication-type="article-journal"><article-title>Tree diversity enhance species richness of beneficial insect in experimental biodiversity enrichment in oil palm plantation</article-title><source>International Journal of Oil Palm</source><volume>5</volume><person-group person-group-type="author"><name><surname>Azhar</surname><given-names>A.</given-names></name><name><surname>Tawakkal</surname><given-names>M.I.</given-names></name><name><surname>Sari</surname><given-names>A.</given-names></name><name><surname>Rizali</surname><given-names>A.</given-names></name><name><surname>Tarigan</surname><given-names>S.D.</given-names></name><name><surname>Nazarreta</surname><given-names>R.</given-names></name><name><surname>Buchori</surname><given-names>D.</given-names></name></person-group><year>2022</year><fpage>39</fpage><lpage>49</lpage><page-range>39-49</page-range><pub-id pub-id-type="doi">10.35876/ijop.v5i2.82</pub-id></element-citation></ref><ref id="BIBR-7"><element-citation publication-type="article-journal"><article-title>Arthropod diversity in a tropical forest</article-title><source>Science</source><volume>338</volume><person-group person-group-type="author"><name><surname>Basset</surname><given-names>Y.</given-names></name><name><surname>Cizek</surname><given-names>L.</given-names></name><name><surname>Cuénoud</surname><given-names>P.</given-names></name><name><surname>Didham</surname><given-names>R.K.</given-names></name><name><surname>Guilhaumon</surname><given-names>F.</given-names></name><name><surname>Missa</surname><given-names>O.</given-names></name><name><surname>Novotny</surname><given-names>V.</given-names></name><name><surname>Ødegaard</surname><given-names>F.</given-names></name><name><surname>Roslin</surname><given-names>T.</given-names></name><name><surname>Schmidl</surname><given-names>J.</given-names></name><name><surname>Tishechkin</surname><given-names>A.K.</given-names></name><name><surname>Winchester</surname><given-names>N.N.</given-names></name><name><surname>Roubik</surname><given-names>D.W.</given-names></name><name><surname>Aberlenc</surname><given-names>H.</given-names></name><name><surname>Bail</surname><given-names>J.</given-names></name><name><surname>Barrios</surname><given-names>H.</given-names></name><name><surname>Bridle</surname><given-names>Castaño-Meneses</given-names></name><name><surname>G</surname><given-names>Corbara</given-names></name><name><surname>B</surname><given-names>Curletti</given-names></name><name><surname>G</surname><given-names>da Rocha</given-names></name><name><surname>WD</surname><given-names>De Bakker</given-names></name><name><surname>D</surname><given-names>Delabie</given-names></name><name><surname>JHC</surname><given-names>Dejean</given-names></name><name><surname>A</surname><given-names>Fagan</given-names></name><name><surname>LL</surname><given-names>Floren</given-names></name><name><surname>A</surname><given-names>Kitching</given-names></name><name><surname>RL</surname><given-names>Medianero</given-names></name><name><surname>E</surname><given-names>Miller</given-names></name><name><surname>SE</surname><given-names>de Oliveira</given-names></name><name><surname>EG</surname><given-names>Orivel</given-names></name><name><surname>J</surname><given-names>Pollet</given-names></name><name><surname>M</surname><given-names>Rapp</given-names></name><name><surname>M</surname><given-names>Ribeiro</given-names></name><name><surname>SP</surname><given-names>Roisin</given-names></name><name><surname>Y</surname><given-names>Schmidt</given-names></name><name><surname>JB</surname><given-names>Sørensen</given-names></name><name><surname>L</surname><given-names>Leponce</given-names></name><name name-style="given-only"><given-names>M.</given-names></name></person-group><year>2012</year><fpage>1481</fpage><lpage>1484</lpage><page-range>1481-1484</page-range><pub-id pub-id-type="doi">10.1126/science.1226727</pub-id></element-citation></ref><ref id="BIBR-8"><element-citation publication-type="article-journal"><article-title>Environmental and evolutionary factors favouring the coexistence of sarcosaprophagous Calliphoridae species competing for animal necromass</article-title><source>Ecological Entomology</source><volume>46</volume><person-group person-group-type="author"><name><surname>Boudreau</surname><given-names>D.R.</given-names></name><name><surname>Hammami</surname><given-names>N.</given-names></name><name><surname>Moreau</surname><given-names>G.</given-names></name></person-group><year>2021</year><fpage>1</fpage><lpage>8</lpage><page-range>1-8</page-range><pub-id pub-id-type="doi">10.1111/een.13076</pub-id></element-citation></ref><ref id="BIBR-9"><element-citation publication-type="journal"><article-title>Luas Kawasan Hutan dan Perairan Menurut Kabupaten/Kota di Provinsi Jambi (ha</article-title><person-group person-group-type="author"><name><surname>Jambi</surname><given-names>B.P.S.] Badan Pusat Statistik Provinsi</given-names></name></person-group><year>2025</year><comment>BPS Provinsi Jambi. Available at:</comment><ext-link xlink:href="https://jambi.bps.go.id/id/statistics-table/3Umpod09GaG5RbTlOYTJwaVFXWnJPRmQ1Wm1wMVp6MDkjMw==/luas-kawasan-hutan-dan-konservasi-perairan-sup--1--sup--menurut-kabupaten-kota-dan-fungsi-hutan-di-provinsi-jambi--ha---2023.html?year=2023[accessed" ext-link-type="uri">https://jambi.bps.go.id/id/statistics-table/3Umpod09GaG5RbTlOYTJwaVFXWnJPRmQ1Wm1wMVp6MDkjMw==/luas-kawasan-hutan-dan-konservasi-perairan-sup--1--sup--menurut-kabupaten-kota-dan-fungsi-hutan-di-provinsi-jambi--ha---2023.html?year=2023[accessed</ext-link></element-citation></ref><ref id="BIBR-10"><element-citation publication-type="book"><article-title>Provinsi Jambi dalam Angka 2025</article-title><person-group person-group-type="author"><name><surname>Jambi</surname><given-names>B.P.S.] Badan Pusat Statistik Provinsi</given-names></name></person-group><year>2025</year><publisher-name>BPS Provinsi Jambi</publisher-name><publisher-loc>Jakarta</publisher-loc></element-citation></ref><ref id="BIBR-11"><element-citation publication-type="article-journal"><article-title>Partitioning the turnover and nestedness components of beta diversity</article-title><source>Global Ecology and Biogeography</source><volume>19</volume><person-group person-group-type="author"><name><surname>Baselga</surname><given-names>A.</given-names></name></person-group><year>2010</year><fpage>134</fpage><lpage>143</lpage><page-range>134-143</page-range><pub-id pub-id-type="doi">10.1111/j.1466-8238.2009.00490.x</pub-id></element-citation></ref><ref id="BIBR-12"><element-citation publication-type="journal"><article-title>Ggboral: view BORAL model results with ggplot</article-title><person-group person-group-type="author"><name><surname>Bedward</surname><given-names>M.</given-names></name></person-group><year>2020</year><comment>Available at: https://github.com/mbedward/ggboral[accessed 13 May 2025</comment></element-citation></ref><ref id="BIBR-13"><element-citation publication-type="article-journal"><article-title>Effects of land-use change on vascular epiphyte diversity in Sumatra (Indonesia</article-title><source>Biological Conservation</source><volume>202</volume><person-group person-group-type="author"><name><surname>Böhnert</surname><given-names>T.</given-names></name><name><surname>Wenzel</surname><given-names>A.</given-names></name><name><surname>Altenhövel</surname><given-names>C.</given-names></name><name><surname>Beeretz</surname><given-names>L.</given-names></name><name><surname>Tjitrosoedirdjo</surname><given-names>S.S.</given-names></name><name><surname>Meijide</surname><given-names>A.</given-names></name><name><surname>Rembold</surname><given-names>K.</given-names></name><name><surname>Kreft</surname><given-names>H.</given-names></name></person-group><year>2016</year><fpage>20</fpage><lpage>29</lpage><page-range>20-29</page-range><pub-id pub-id-type="doi">10.1016/j.biocon.2016.08.008</pub-id></element-citation></ref><ref id="BIBR-14"><element-citation publication-type="article-journal"><article-title>Seasonal yeast compositions in Forcipomyia taiwana (Diptera: Ceratopogonidae</article-title><source>Journal Asia-Pacific Entomology</source><volume>19</volume><person-group person-group-type="author"><name><surname>Chen</surname><given-names>H.</given-names></name><name><surname>Chou</surname><given-names>J.</given-names></name><name><surname>Lin</surname><given-names>C.</given-names></name><name><surname>Wen</surname><given-names>Y.</given-names></name><name><surname>Wang</surname><given-names>W.</given-names></name></person-group><year>2016</year><fpage>509</fpage><lpage>514</lpage><page-range>509-514</page-range><pub-id pub-id-type="doi">10.1016/j.aspen.2016.04.020</pub-id></element-citation></ref><ref id="BIBR-15"><element-citation publication-type="article-journal"><article-title>Dampak konversi hutan menjadi perkebunan kelapa sawit terhadap keanekaragaman dan kelimpahan kupu-kupu Superfamili Papilionoidea</article-title><source>Jurnal Biologi Papua</source><volume>15</volume><person-group person-group-type="author"><name><surname>Daawia</surname><given-names>D.</given-names></name><name><surname>Dianingsih</surname><given-names>N.</given-names></name></person-group><year>2023</year><fpage>28</fpage><lpage>38</lpage><page-range>28-38</page-range><pub-id pub-id-type="doi">10.31957/jbp.2680</pub-id></element-citation></ref><ref id="BIBR-16"><element-citation publication-type="article-journal"><article-title>Vertical stratification of insect abundance and species richness in an Amazonian tropical forest</article-title><source>Scientific Reports</source><volume>12</volume><issue>1734</issue><person-group person-group-type="author"><string-name>de Souza Amorim D, Brown BV, Boscolo D. Ale-Rocha R, Alvarez-Garcia DM, Balbi MIPA, Barbosa AD, Capellari RS, de Carvalho CJB, Couri MS, Dioz RDP, Fachin DA, Ferro GB, Flores HF, Frare LM, Gudin FM, Hauser M, Lamas CJE, Lindsay KG, Marinho MAT, Marques DWA, Marshall SA, Mello-Patiu C, Menezes MA, Morales MN, Nihei SS, Oliveira SS, Pirani G, Ribeiro GC, Riccardi PR, de Santis MD, Santos D, dos Santos JR, Silva VC, Wood EM, Rafael JA</string-name></person-group><year>2022</year><pub-id pub-id-type="doi">10.1038/s41598-022-05677-y</pub-id></element-citation></ref><ref id="BIBR-17"><element-citation publication-type="article-journal"><source>IOP Conference Series: Earth and Environmental Science</source><volume>1220</volume><issue>012025</issue><person-group person-group-type="author"><name><surname>Desriana</surname><given-names>R.</given-names></name><name><surname>Buchori</surname><given-names>D.</given-names></name><name><surname>Maryana</surname><given-names>N.</given-names></name></person-group><year>2023</year><pub-id pub-id-type="doi">10.1088/1755-1315/1220/1/012025</pub-id></element-citation></ref><ref id="BIBR-18"><element-citation publication-type="article-journal"><article-title>Ecological and socio-economic functions across tropical land use systems after rainforest conversion</article-title><source>Philosophical Transactions of the Royal Society B: Biological Sciences</source><volume>371</volume><person-group person-group-type="author"><name><surname>Drescher</surname><given-names>J.</given-names></name><name><surname>Rembold</surname><given-names>K.</given-names></name><name><surname>Allen</surname><given-names>K.</given-names></name><name><surname>Beckschäfer</surname><given-names>P.</given-names></name><name><surname>Buchori</surname><given-names>D.</given-names></name><name><surname>Clough</surname><given-names>Y.</given-names></name><name><surname>Faust</surname><given-names>H.</given-names></name><name><surname>Fauzi</surname><given-names>A.M.</given-names></name><name><surname>Gunawan</surname><given-names>D.</given-names></name><name><surname>Hertel</surname><given-names>D.</given-names></name><name><surname>Irawan</surname><given-names>B.</given-names></name><name><surname>Jaya</surname><given-names>N.S.</given-names></name><name><surname>Klarner</surname><given-names>B.</given-names></name><name><surname>Kleinn</surname><given-names>C.</given-names></name><name><surname>Knohl</surname><given-names>A.</given-names></name><name><surname>Kotowska</surname><given-names>M.M.</given-names></name><name><surname>Krashevska</surname><given-names>V.</given-names></name><name><surname>Krishna</surname><given-names>V.</given-names></name><name><surname>Leuschner</surname><given-names>C.</given-names></name><name><surname>Lorenz</surname><given-names>W.</given-names></name><name><surname>Meijide</surname><given-names>A.</given-names></name><name><surname>Melati</surname><given-names>D.</given-names></name><name><surname>Nomura</surname><given-names>N.</given-names></name><name><surname>Perez-Cruzado</surname><given-names>C.</given-names></name><name><surname>Qaim</surname><given-names>M.</given-names></name><name><surname>Siregar</surname><given-names>I.Z.</given-names></name><name><surname>Steinebach</surname><given-names>S.</given-names></name><name><surname>Tjoa</surname><given-names>A.</given-names></name><name><surname>Tscharntke</surname><given-names>T.</given-names></name><name><surname>Wick</surname><given-names>B.</given-names></name><name><surname>Wiegand</surname><given-names>K.</given-names></name><name><surname>Kreft</surname><given-names>H.</given-names></name><name><surname>Scheu</surname><given-names>S.</given-names></name></person-group><year>2016</year><fpage>1</fpage><lpage>8</lpage><page-range>1-8</page-range><pub-id pub-id-type="doi">10.1098/rstb.2015.0275</pub-id></element-citation></ref><ref id="BIBR-19"><element-citation publication-type="article-journal"><article-title>Trophic change and community decline in acrobat ants after rainforest conversion to cash crops</article-title><source>Ecology and Evolution</source><volume>14</volume><person-group person-group-type="author"><name><surname>TR</surname><given-names>Ehlers J.Hartke</given-names></name><name><surname>N</surname><given-names>Janotta</given-names></name><name><surname>A</surname><given-names>Mawan</given-names></name><name><surname>R</surname><given-names>Nazarreta</given-names></name><name><surname>R</surname><given-names>Desriana</given-names></name><name><surname>P</surname><given-names>Hidayat</given-names></name><name><surname>D</surname><given-names>Buchori</given-names></name><name><surname>S</surname><given-names>Scheu</given-names></name><name><surname>MM</surname><given-names>Pollierer</given-names></name><name><surname>J</surname><given-names>Drescher</given-names></name></person-group><year>2024</year><fpage>1</fpage><lpage>11</lpage><page-range>1-11</page-range><pub-id pub-id-type="doi">10.1002/ece3.70694</pub-id></element-citation></ref><ref id="BIBR-20"><element-citation publication-type="article-journal"><article-title>Oviposition of Culicoides insignis (Diptera: Ceratopogonidae) under laboratory conditions with notes on the developmental life history traits of its immature stages</article-title><source>Parasit Vectors</source><volume>14</volume><issue>1</issue><person-group person-group-type="author"><name><surname>Erram</surname><given-names>D.</given-names></name><name><surname>Burkett-Cadena</surname><given-names>N.</given-names></name></person-group><year>2021</year><page-range>9</page-range><comment>DOI:</comment><pub-id pub-id-type="doi">10.1186/s13071-021-05025-5</pub-id></element-citation></ref><ref id="BIBR-21"><element-citation publication-type="article-journal"><article-title>Why do calling medflies (Diptera: Tephritidae) cluster? Assessing the empirical evidence for models of medfly lek evolution</article-title><source>Florida Entomologist</source><volume>85</volume><person-group person-group-type="author"><name><surname>Field</surname><given-names>S.A.</given-names></name><name><surname>Kaspi</surname><given-names>R.</given-names></name><name><surname>Yuval</surname><given-names>B.</given-names></name></person-group><year>2003</year><fpage>63</fpage><lpage>72</lpage><page-range>63-72</page-range><pub-id pub-id-type="doi">10.1653/0015-4040(2002)085</pub-id></element-citation></ref><ref id="BIBR-22"><element-citation publication-type="article-journal"><article-title>Taxonomic, species and functional group diversity of ants in a tropical anthropogenic landscape</article-title><source>Tropical Conservation Science</source><volume>8</volume><person-group person-group-type="author"><name><surname>M.Á</surname><given-names>García-Martínez</given-names></name><name><surname>DL</surname><given-names>Martínez-Tlapa</given-names></name><name><surname>G.R</surname><given-names>Pérez-Toledo</given-names></name><name><surname>LN</surname><given-names>Quiroz-Robledo</given-names></name><name><surname>G</surname><given-names>Castaño-Meneses</given-names></name><name><surname>J</surname><given-names>Laborde</given-names></name><name><surname>JE</surname><given-names>Valenzuela-González</given-names></name></person-group><year>2015</year><fpage>1017</fpage><lpage>1032</lpage><page-range>1017-1032</page-range><pub-id pub-id-type="doi">10.1177/194008291500800412</pub-id></element-citation></ref><ref id="BIBR-23"><element-citation publication-type="journal"><article-title>Global Biodiversity Information Facility</article-title><person-group person-group-type="author"><name name-style="given-only"><given-names>G.B.I.F.</given-names></name></person-group><year>2025</year><comment>Diptera. Available at:</comment><ext-link xlink:href="https://www.gbif.org/" ext-link-type="uri">https://www.gbif.org/</ext-link></element-citation></ref><ref id="BIBR-24"><element-citation publication-type="article-journal"><article-title>Intensification of Ethiopian coffee agroforestry drives impoverishment of the Arabica coffee flower visiting bee and fly communities</article-title><source>Agroforestry Systems</source><volume>93</volume><person-group person-group-type="author"><name><surname>Geeraert</surname><given-names>L.</given-names></name><name><surname>Aerts</surname><given-names>R.</given-names></name><name><surname>Jordaens</surname><given-names>K.</given-names></name><name><surname>Dox</surname><given-names>I.</given-names></name><name><surname>Wellens</surname><given-names>S.</given-names></name><name><surname>Couri</surname><given-names>M.</given-names></name><name><surname>Berecha</surname><given-names>G.</given-names></name><name><surname>Honnay</surname><given-names>O.</given-names></name></person-group><year>2018</year><fpage>1729</fpage><lpage>1739</lpage><page-range>1729-1739</page-range><pub-id pub-id-type="doi">10.1007/s10457-018-0280-0</pub-id></element-citation></ref><ref id="BIBR-25"><element-citation publication-type="article-journal"><article-title>Oil-palm yields in diversified plantations: initial results from a biodiversity enrichment experiment in Sumatra, Indonesia</article-title><source>Agriculture, Ecosystems and Environment</source><volume>240</volume><person-group person-group-type="author"><name><surname>Gérard</surname><given-names>A.</given-names></name><name><surname>Wollni</surname><given-names>M.</given-names></name><name><surname>Hölscher</surname><given-names>D.</given-names></name><name><surname>Irawan</surname><given-names>B.</given-names></name><name><surname>Sundawati</surname><given-names>L.</given-names></name><name><surname>Teuscher</surname><given-names>M.</given-names></name><name><surname>Kreft</surname><given-names>H.</given-names></name></person-group><year>2017</year><fpage>253</fpage><lpage>260</lpage><page-range>253-260</page-range><comment>DOI:</comment><pub-id pub-id-type="doi">10.1016/j.agee.2017.02.026</pub-id></element-citation></ref><ref id="BIBR-26"><element-citation publication-type="article-journal"><article-title>Primary forests are irreplaceable for sustaining tropical biodiversity</article-title><source>Nature</source><volume>478</volume><person-group person-group-type="author"><name><surname>Gibson</surname><given-names>L.</given-names></name><name><surname>Lee</surname><given-names>T.M.</given-names></name><name><surname>Koh</surname><given-names>L.P.</given-names></name><name><surname>Brook</surname><given-names>B.W.</given-names></name><name><surname>Gardner</surname><given-names>T.A.</given-names></name><name><surname>Barlow</surname><given-names>J.</given-names></name><name><surname>Peres</surname><given-names>C.A.</given-names></name><name><surname>Bradshaw</surname><given-names>C.J.A.</given-names></name><name><surname>Laurance</surname><given-names>W.F.</given-names></name><name><surname>Lovejoy</surname><given-names>T.E.</given-names></name><name><surname>Sodhi</surname><given-names>N.S.</given-names></name></person-group><year>2011</year><fpage>378</fpage><lpage>381</lpage><page-range>378-381</page-range><pub-id pub-id-type="doi">10.1038/nature10425</pub-id></element-citation></ref><ref id="BIBR-27"><element-citation publication-type="article-journal"><article-title>Trade-offs between multifunctionality and profit in tropical smallholder landscapes</article-title><source>Nature Communications</source><volume>11</volume><person-group person-group-type="author"><name><surname>Grass</surname><given-names>I.</given-names></name><name><surname>Kubitza</surname><given-names>C.</given-names></name><name><surname>V.</surname><given-names>Krishna V.</given-names></name><name><surname>Corre</surname><given-names>M.D.</given-names></name><name><surname>Mußhoff</surname><given-names>O.</given-names></name><name><surname>Pütz</surname><given-names>P.</given-names></name><name><surname>Drescher</surname><given-names>J.</given-names></name><name><surname>Rembold</surname><given-names>K.</given-names></name><name><surname>Ariyanti</surname><given-names>E.S.</given-names></name><name><surname>Barnes</surname><given-names>A.D.</given-names></name><name><surname>Brinkmann</surname><given-names>N.</given-names></name><name><surname>Brose</surname><given-names>U.</given-names></name><name><surname>Brümmer</surname><given-names>B.</given-names></name><name><surname>Buchori</surname><given-names>D.</given-names></name><name><surname>Daniel</surname><given-names>R.</given-names></name><name><surname>Darras</surname><given-names>K.A.</given-names></name><name><surname>Faust</surname><given-names>H.</given-names></name><name><surname>Fehrmann</surname><given-names>L.</given-names></name><name><surname>Hein</surname><given-names>J.</given-names></name><name><surname>Hennings</surname><given-names>N.</given-names></name><name><surname>Hidayat</surname><given-names>P.</given-names></name><name><surname>Hölscher</surname><given-names>D.</given-names></name><name><surname>Jochum</surname><given-names>M.</given-names></name><name><surname>Knohl</surname><given-names>A.</given-names></name><name><surname>Kotowska</surname><given-names>M.M.</given-names></name><name><surname>Krashevska</surname><given-names>V.</given-names></name><name><surname>Kreft</surname><given-names>H.</given-names></name><name><surname>Leuschner</surname><given-names>C.</given-names></name><name><surname>Lobite</surname><given-names>N.J.S.</given-names></name><name><surname>Panjaitan</surname><given-names>R.</given-names></name><name><surname>Polle</surname><given-names>A.</given-names></name><name><surname>Potapov</surname><given-names>A.M.</given-names></name><name><surname>Purnama</surname><given-names>E.</given-names></name><name><surname>Qaim</surname><given-names>M.</given-names></name><name><surname>Röll</surname><given-names>A.</given-names></name><name><surname>Scheu</surname><given-names>S.</given-names></name><name><surname>Schneider</surname><given-names>D.</given-names></name><name><surname>Tjoa</surname><given-names>A.</given-names></name><name><surname>Tscharntke</surname><given-names>T.</given-names></name><name><surname>Veldkamp</surname><given-names>E.</given-names></name><name><surname>Wollni</surname><given-names>M.</given-names></name></person-group><year>2020</year><fpage>1</fpage><lpage>13</lpage><page-range>1-13</page-range><pub-id pub-id-type="doi">10.1038/s41467-020-15013-5</pub-id></element-citation></ref><ref id="BIBR-28"><element-citation publication-type="article-journal"><article-title>Komunitas semut (Hymenoptera: Formicidae) pada empat tipe ekosistem yang berbeda di Desa Bungku Provinsi Jambi</article-title><source>Jurnal Silvikultur Tropika</source><volume>6</volume><person-group person-group-type="author"><name><surname>Haneda</surname><given-names>N.F.</given-names></name><name><surname>Yuniar</surname><given-names>N.</given-names></name></person-group><year>2015</year><fpage>203</fpage><lpage>209</lpage><page-range>203-209</page-range><pub-id pub-id-type="doi">10.13057/psnmbi/m010707</pub-id></element-citation></ref><ref id="BIBR-29"><element-citation publication-type="journal"><article-title>RankAbund: calculate rank abundances and plot rank abundance curves</article-title><person-group person-group-type="author"><name><surname>Hartke</surname><given-names>T.</given-names></name></person-group><year>2020</year><comment>Available at: https://github.com/tamarahartke/RankAbund[accessed 13 May 2025</comment></element-citation></ref><ref id="BIBR-30"><element-citation publication-type="article-journal"><article-title>Bleeding’ flowers of Ceropegia gerrardii (Apocynaceae-Asclepiadoideae) mimic wounded insects to attract kleptoparasitic fly pollinators</article-title><source>New Phytologist</source><volume>239</volume><person-group person-group-type="author"><name><surname>Heiduk</surname><given-names>A.</given-names></name><name><surname>Brake</surname><given-names>I.</given-names></name><name><surname>Shuttleworth</surname><given-names>A.</given-names></name><name><surname>Johnson</surname><given-names>S.D.</given-names></name></person-group><year>2023</year><fpage>1490</fpage><lpage>1504</lpage><page-range>1490-1504</page-range><pub-id pub-id-type="doi">10.1111/nph.18888</pub-id></element-citation></ref><ref id="BIBR-31"><element-citation publication-type="journal"><article-title>Boral: bayesian ordination and regression analysis</article-title><person-group person-group-type="author"><name><surname>Hui</surname><given-names>F.K.C.</given-names></name></person-group><year>2025</year></element-citation></ref><ref id="BIBR-32"><element-citation publication-type="article-journal"><article-title>Food web structure affects the extinction risk of species in ecological communities</article-title><source>Handbook of Environmental Chemistry</source><volume>5</volume><person-group person-group-type="author"><name><surname>Jonsson</surname><given-names>T.</given-names></name><name><surname>Karlsson</surname><given-names>P.</given-names></name><name><surname>Jonsson</surname><given-names>A.</given-names></name></person-group><year>2006</year><fpage>93</fpage><lpage>106</lpage><page-range>93-106</page-range><pub-id pub-id-type="doi">10.1016/j.ecolmodel.2006.06.012</pub-id></element-citation></ref><ref id="BIBR-33"><element-citation publication-type="article-journal"><article-title>Changes in diversity and community assembly of jumping spiders (Araneae: Salticidae) after rainforest conversion to rubber and oil palm plantations</article-title><source>PeerJ</source><volume>9</volume><person-group person-group-type="author"><name><surname>Junggebauer</surname><given-names>A.</given-names></name><name><surname>Hartke</surname><given-names>T.R.</given-names></name><name><surname>Ramos</surname><given-names>D.</given-names></name><name><surname>Schaefer</surname><given-names>I.</given-names></name><name><surname>Buchori</surname><given-names>D.</given-names></name><name><surname>Hidayat</surname><given-names>P.</given-names></name><name><surname>Scheu</surname><given-names>S.</given-names></name><name><surname>Drescher</surname><given-names>J.</given-names></name></person-group><year>2021</year><fpage>1</fpage><lpage>26</lpage><page-range>1-26</page-range><pub-id pub-id-type="doi">10.7717/peerj.11012</pub-id></element-citation></ref><ref id="BIBR-34"><element-citation publication-type="article-journal"><article-title>Rainforest conversion to smallholder cash crops leads to varying declines of beetles (Coleoptera) on Sumatra</article-title><source>Biotropica</source><volume>55</volume><person-group person-group-type="author"><name><surname>Kasmiatun</surname><given-names>Hartke T.R.</given-names></name><name><surname>D</surname><given-names>Buchori</given-names></name><name><surname>P</surname><given-names>Hidayat</given-names></name><name><surname>F</surname><given-names>Siddikah</given-names></name><name><surname>R</surname><given-names>Amrulloh</given-names></name><name><surname>MS</surname><given-names>Hiola</given-names></name><name><surname>L</surname><given-names>Najmi</given-names></name><name><surname>WA</surname><given-names>Noerdjito</given-names></name><name><surname>S</surname><given-names>Scheu</given-names></name><name><surname>J</surname><given-names>Drescher</given-names></name></person-group><year>2022</year><fpage>119</fpage><lpage>131</lpage><page-range>119-131</page-range><pub-id pub-id-type="doi">10.1111/btp.13165</pub-id></element-citation></ref><ref id="BIBR-35"><element-citation publication-type="article-journal"><article-title>Mediterranean fruit fly leks: factors affecting male location</article-title><source>Functional Ecology</source><volume>13</volume><person-group person-group-type="author"><name><surname>Kaspi</surname><given-names>R.</given-names></name><name><surname>Yuval</surname><given-names>B.</given-names></name></person-group><year>1999</year><fpage>539</fpage><lpage>545</lpage><page-range>539-545</page-range><pub-id pub-id-type="doi">10.1046/j.1365-2435.1999.00344.x</pub-id></element-citation></ref><ref id="BIBR-36"><element-citation publication-type="book"><article-title>Manual of Afrotropical Diptera</article-title><volume>1</volume><person-group person-group-type="author"><name><surname>Kirk-Spriggs</surname><given-names>A.H.</given-names></name><name><surname>Sinclair</surname><given-names>B.J.</given-names></name><name><surname>Marshall</surname><given-names>S.A.</given-names></name></person-group><year>2017</year><publisher-name>South African National Biodiversity Institute (SANBI Publishing</publisher-name><publisher-loc>Pretoria</publisher-loc></element-citation></ref><ref id="BIBR-37"><element-citation publication-type="book"><article-title>Manual of Afrotropical Diptera</article-title><volume>2</volume><person-group person-group-type="author"><name><surname>Kirk-Spriggs</surname><given-names>A.H.</given-names></name><name><surname>Sinclair</surname><given-names>B.J.</given-names></name><name><surname>Marshall</surname><given-names>S.A.</given-names></name></person-group><year>2017</year><publisher-name>South African National Biodiversity Institute (SANBI Publishing</publisher-name><publisher-loc>Pretoria</publisher-loc></element-citation></ref><ref id="BIBR-38"><element-citation publication-type="book"><article-title>Manual of Afrotropical Diptera</article-title><volume>3</volume><person-group person-group-type="author"><name><surname>Kirk-Spriggs</surname><given-names>A.H.</given-names></name><name><surname>Sinclair</surname><given-names>B.J.</given-names></name><name><surname>Marshall</surname><given-names>S.A.</given-names></name></person-group><year>2021</year><publisher-name>South African National Biodiversity Institute (SANBI Publishing</publisher-name><publisher-loc>Pretoria</publisher-loc></element-citation></ref><ref id="BIBR-39"><element-citation publication-type="article-journal"><article-title>Eco-floristic sectors and deforestation threats in Sumatra: identifying new conservation area network priorities for ecosystem-based land use planning</article-title><source>Biodiversity and Conservation</source><volume>19</volume><person-group person-group-type="author"><name><surname>Laumonier</surname><given-names>Y.</given-names></name><name><surname>Uryu</surname><given-names>Y.</given-names></name><name><surname>Stüwe</surname><given-names>M.</given-names></name><name><surname>Budiman</surname><given-names>A.</given-names></name><name><surname>Setiabudi</surname><given-names>B.</given-names></name><name><surname>Hadian</surname><given-names>O.</given-names></name></person-group><year>2010</year><fpage>1153</fpage><lpage>1174</lpage><page-range>1153-1174</page-range><pub-id pub-id-type="doi">10.1007/s10531-010-9784-2</pub-id></element-citation></ref><ref id="BIBR-40"><element-citation publication-type="article-journal"><article-title>Five new species of Asphondylia (Diptera, Cecidomyiidae, Asphondyliini) from Brazilian restinga (Atlantic Forest</article-title><source>Papéis Avulsos de Zoologia</source><volume>64</volume><person-group person-group-type="author"><name><surname>Maia</surname><given-names>V.C.</given-names></name></person-group><year>2024</year><fpage>1</fpage><lpage>30</lpage><page-range>1-30</page-range><pub-id pub-id-type="doi">10.11606/1807-0205/2024.64.008</pub-id></element-citation></ref><ref id="BIBR-41"><element-citation publication-type="article-journal"><article-title>Primary forest cover loss in Indonesia over 2000 – 2012</article-title><source>Nature Climate Change</source><volume>4</volume><person-group person-group-type="author"><name><surname>Margono</surname><given-names>B.A.</given-names></name><name><surname>Potapov</surname><given-names>P.V.</given-names></name><name><surname>Turubanova</surname><given-names>S.</given-names></name><name><surname>Stolle</surname><given-names>F.</given-names></name><name><surname>Hansen</surname><given-names>M.C.</given-names></name></person-group><year>2014</year><fpage>730</fpage><lpage>735</lpage><page-range>730-735</page-range><pub-id pub-id-type="doi">10.1038/nclimate2277</pub-id></element-citation></ref><ref id="BIBR-42"><element-citation publication-type="book"><article-title>Manual of Nearctic Diptera</article-title><volume>1</volume><person-group person-group-type="author"><name><surname>McAlpine</surname><given-names>J.F.</given-names></name><name><surname>Peterson</surname><given-names>B.V.</given-names></name><name><surname>Shewell</surname><given-names>G.E.</given-names></name><name><surname>Teskey</surname><given-names>H.J.</given-names></name><name><surname>Vockeroth</surname><given-names>Wood</given-names></name><name name-style="given-only"><given-names>D.M.</given-names></name></person-group><year>1981</year><publisher-name>Biosystematics Research Institute</publisher-name><publisher-loc>Ottawa</publisher-loc></element-citation></ref><ref id="BIBR-43"><element-citation publication-type="book"><article-title>Manual of Nearctic Diptera</article-title><volume>2</volume><person-group person-group-type="author"><name><surname>McAlpine</surname><given-names>J.F.</given-names></name><name><surname>Peterson</surname><given-names>B.V.</given-names></name><name><surname>Shewell</surname><given-names>G.E.</given-names></name><name><surname>Teskey</surname><given-names>H.J.</given-names></name><name><surname>Vockeroth</surname><given-names>Wood</given-names></name><name name-style="given-only"><given-names>D.M.</given-names></name></person-group><year>1987</year><publisher-name>Research Branch</publisher-name><publisher-loc>Ottawa</publisher-loc></element-citation></ref><ref id="BIBR-44"><element-citation publication-type="article-journal"><article-title>The filtering effect of oil palm plantation on potential insect pollinator assemblages from remnant forest patches</article-title><source>Land</source><volume>12</volume><person-group person-group-type="author"><name><surname>Mohd-Azlan</surname><given-names>J.</given-names></name><name><surname>Conway</surname><given-names>S.</given-names></name><name><surname>Travers</surname><given-names>T.J.</given-names></name><name><surname>Lawes</surname><given-names>M.J.</given-names></name></person-group><year>2023</year><fpage>1</fpage><lpage>20</lpage><page-range>1-20</page-range><pub-id pub-id-type="doi">10.3390/land12061256</pub-id></element-citation></ref><ref id="BIBR-45"><element-citation publication-type="article-journal"><article-title>Biodiversity hotspots for conservation priorities</article-title><source>Nature</source><volume>403</volume><person-group person-group-type="author"><name><surname>Myers</surname><given-names>N.</given-names></name><name><surname>Mittermeier</surname><given-names>R.A.</given-names></name><name><surname>Mitterleier</surname><given-names>C.G.</given-names></name><name><surname>Fonseca</surname><given-names>G.A.B.</given-names></name><name><surname>Kent</surname><given-names>J.</given-names></name></person-group><year>2000</year><fpage>853</fpage><lpage>858</lpage><page-range>853-858</page-range><pub-id pub-id-type="doi">10.1038/35002501</pub-id></element-citation></ref><ref id="BIBR-46"><element-citation publication-type="article-journal"><article-title>Vertebrate blood-feeding biting midges of the Subgenus Lasiohelea Kieffer of Forcipomyia Meigen in Europe (Diptera: Ceratopogonidae) with new synonyms</article-title><source>Annales Zoologici</source><volume>67</volume><person-group person-group-type="author"><name><surname>Navai</surname><given-names>S.</given-names></name><name><surname>Dominiak</surname><given-names>P.</given-names></name><name><surname>Szadziewski</surname><given-names>R.</given-names></name></person-group><year>2017</year><fpage>823</fpage><lpage>835</lpage><page-range>823-835</page-range><pub-id pub-id-type="doi">10.3161/00034541ANZ2017.67.4.016</pub-id></element-citation></ref><ref id="BIBR-47"><element-citation publication-type="article-journal"><article-title>Rainforest conversion to smallholder plantations of rubber or oil palm leads to species loss and community shifts in canopy ants (Hymenoptera: Formicidae</article-title><source>Myrmecological News</source><volume>30</volume><person-group person-group-type="author"><name><surname>Nazarreta</surname><given-names>R.</given-names></name><name><surname>Hartke</surname><given-names>T.R.</given-names></name><name><surname>Hidayat</surname><given-names>P.</given-names></name><name><surname>Scheu</surname><given-names>S.</given-names></name><name><surname>Buchori</surname><given-names>D.</given-names></name><name><surname>Drescher</surname><given-names>J.</given-names></name></person-group><year>2020</year><fpage>175</fpage><lpage>186</lpage><page-range>175-186</page-range><pub-id pub-id-type="doi">10.25849/myrmecol.news_030175</pub-id></element-citation></ref><ref id="BIBR-48"><element-citation publication-type="article-journal"><article-title>Pembaruan informasi taksonomi nyamuk dan kunci identifikasi fotografis genus nyamuk (Diptera: Culicidae) di Indonesia</article-title><source>Jurnal Entomologi Indonesia</source><volume>18</volume><person-group person-group-type="author"><name><surname>Nugroho</surname><given-names>S.S.</given-names></name><name><surname>Mujiyono</surname><given-names>M.</given-names></name></person-group><year>2021</year><fpage>55</fpage><lpage>71</lpage><page-range>55-71</page-range><pub-id pub-id-type="doi">10.5994/jei.18.1.55</pub-id></element-citation></ref><ref id="BIBR-49"><element-citation publication-type="journal"><article-title>Vegan: community ecology package</article-title><person-group person-group-type="author"><name><surname>Oksanen</surname><given-names>J.</given-names></name><name><surname>Simpson</surname><given-names>G.</given-names></name><name><surname>Blanchet</surname><given-names>F.</given-names></name><name><surname>Kindt</surname><given-names>R.</given-names></name><name><surname>Legendre</surname><given-names>P.</given-names></name><name><surname>Minchin</surname><given-names>P.</given-names></name><name><surname>O’Hara</surname><given-names>R.</given-names></name><name><surname>Solymos</surname><given-names>P.</given-names></name><name><surname>Stevens</surname><given-names>M.</given-names></name><name><surname>Szoecs</surname><given-names>E.</given-names></name><name><surname>Wagner</surname><given-names>H.</given-names></name><name><surname>Barbour</surname><given-names>M.</given-names></name><name><surname>Bedward</surname><given-names>M.</given-names></name><name><surname>Bolker</surname><given-names>B.</given-names></name><name><surname>Borcard</surname><given-names>D.</given-names></name><name><surname>Carvalho</surname><given-names>G.</given-names></name><name><surname>Chirico</surname><given-names>M.</given-names></name><name><surname>Caceres</surname><given-names>M.</given-names></name><name><surname>Durand</surname><given-names>S.</given-names></name><name><surname>Evangelista</surname><given-names>H.</given-names></name><name><surname>FitzJohn</surname><given-names>R.</given-names></name><name><surname>Friendly</surname><given-names>M.</given-names></name><name><surname>Furneaux</surname><given-names>B.</given-names></name><name><surname>Hannigan</surname><given-names>G.</given-names></name><name><surname>Hill</surname><given-names>M.</given-names></name><name><surname>Lahti</surname><given-names>L.</given-names></name><name><surname>McGlinn</surname><given-names>D.</given-names></name><name><surname>Ouellette</surname><given-names>M.</given-names></name><name><surname>Ribeiro Cunha</surname><given-names>E.</given-names></name><name><surname>Smith</surname><given-names>T.</given-names></name><name><surname>Stier</surname><given-names>A.</given-names></name><name><surname>Ter Braak</surname><given-names>C.</given-names></name><name><surname>Weedon</surname><given-names>J.</given-names></name><name><surname>Borman</surname><given-names>T.</given-names></name></person-group><year>2025</year><pub-id pub-id-type="doi">10.32614/CRAN.package.vegan</pub-id></element-citation></ref><ref id="BIBR-50"><element-citation publication-type="article-journal"><article-title>Diversity of butterflies (Lepidoptera) across rainforest transformation systems in Jambi, Sumatra, Indonesia</article-title><source>Biodiversitas</source><volume>21</volume><person-group person-group-type="author"><name><surname>Panjaitan</surname><given-names>R.</given-names></name><name><surname>Drescher</surname><given-names>J.</given-names></name><name><surname>Buchori</surname><given-names>D.</given-names></name><name><surname>Peggie</surname><given-names>D.</given-names></name><name><surname>Harahap</surname><given-names>I.S.</given-names></name><name><surname>Scheu</surname><given-names>S.</given-names></name><name><surname>Hidayat</surname><given-names>P.</given-names></name></person-group><year>2020</year><fpage>5119</fpage><lpage>5127</lpage><page-range>5119-5127</page-range><pub-id pub-id-type="doi">10.13057/biodiv/d211117</pub-id></element-citation></ref><ref id="BIBR-51"><element-citation publication-type="journal"><article-title>Global Biodiversity Information Facility</article-title><person-group person-group-type="author"><name name-style="given-only"><given-names>G.B.I.F.</given-names></name></person-group><year>2025</year><comment>Diptera. Available at:</comment><ext-link xlink:href="https://www.gbif.org/" ext-link-type="uri">https://www.gbif.org/</ext-link></element-citation></ref><ref id="BIBR-52"><element-citation publication-type="article-journal"><article-title>Rainforest conversion to plantations fundamentally alters energy fluxes and functions in canopy arthropod food webs</article-title><source>Ecology Letters</source><volume>26</volume><person-group person-group-type="author"><name><surname>Pollierer</surname><given-names>M.</given-names></name><name><surname>Drescher</surname><given-names>J.</given-names></name><name><surname>Potapov</surname><given-names>A.</given-names></name><name><surname>Kasmiatun</surname><given-names>K.</given-names></name><name><surname>Mutiari</surname><given-names>M.</given-names></name><name><surname>Nazarreta</surname><given-names>R.</given-names></name><name><surname>Hidayat</surname><given-names>P.</given-names></name><name><surname>Buchori</surname><given-names>D.</given-names></name><name><surname>Scheu</surname><given-names>S.</given-names></name></person-group><year>2023</year><fpage>1663</fpage><lpage>1675</lpage><page-range>1663-1675</page-range><pub-id pub-id-type="doi">10.1111/ele.14276</pub-id></element-citation></ref><ref id="BIBR-53"><element-citation publication-type="article-journal"><article-title>Rainforest transformation reallocates energy from green to brown food webs</article-title><source>Nature</source><volume>627</volume><person-group person-group-type="author"><name><surname>Potapov</surname><given-names>A.M.</given-names></name><name><surname>Drescher</surname><given-names>J.</given-names></name><name><surname>Darras</surname><given-names>K.</given-names></name><name><surname>Wenzel</surname><given-names>A.</given-names></name><name><surname>Janotta</surname><given-names>N.</given-names></name><name><surname>Nazarreta</surname><given-names>R.</given-names></name><name><surname>Kasmiatun</surname><given-names>Laurent</given-names></name><name><surname>V</surname><given-names>Mawan</given-names></name><name><surname>A</surname><given-names>Utari</given-names></name><name><surname>EH</surname><given-names>Pollierer</given-names></name><name><surname>MM</surname><given-names>Rembold</given-names></name><name><surname>K</surname><given-names>Widyastuti</given-names></name><name><surname>R</surname><given-names>Buchori</given-names></name><name><surname>D</surname><given-names>Hidayat</given-names></name><name><surname>P</surname><given-names>Turner</given-names></name><name><surname>E</surname><given-names>Grass</given-names></name><name><surname>I</surname><given-names>Westphal</given-names></name><name><surname>C</surname><given-names>Tscharntke</given-names></name><name><surname>T</surname><given-names>Scheu</given-names></name><name name-style="given-only"><given-names>S.</given-names></name></person-group><year>2024</year><fpage>116</fpage><lpage>122</lpage><page-range>116-122</page-range><pub-id pub-id-type="doi">10.1038/s41586-024-07083-y</pub-id></element-citation></ref><ref id="BIBR-54"><element-citation publication-type="journal"><article-title>Insect Ecology: Behavior, Populations and Communities</article-title><person-group person-group-type="author"><name><surname>Prince</surname><given-names>P.W.</given-names></name><name><surname>Denno</surname><given-names>R.F.</given-names></name><name><surname>Eubanks</surname><given-names>M.D.</given-names></name><name><surname>Finke</surname><given-names>D.L.</given-names></name><name><surname>Kaplan</surname><given-names>I.</given-names></name></person-group><year>2011</year><publisher-name>Cambridge University Press</publisher-name><publisher-loc>New York</publisher-loc><comment>DOI:</comment><pub-id pub-id-type="doi">10.1017/CBO9780511975387</pub-id></element-citation></ref><ref id="BIBR-55"><element-citation publication-type="article-journal"><article-title>Beta diversity of angiosperms in temperate floras of eastern Asia and eastern North America</article-title><source>Ecology Letter</source><volume>8</volume><person-group person-group-type="author"><name><surname>Qian</surname><given-names>H.</given-names></name><name><surname>Ricklefs</surname><given-names>R.E.</given-names></name><name><surname>White</surname><given-names>P.S.</given-names></name></person-group><year>2005</year><fpage>15</fpage><lpage>22</lpage><page-range>15-22</page-range><pub-id pub-id-type="doi">10.1111/j.1461-0248.2004.00682.x.</pub-id></element-citation></ref><ref id="BIBR-56"><element-citation publication-type="article-journal"><article-title>Tracking community timing: pattern and determinants of seasonality in Culicoides (Diptera: Ceratopogonidae) in Northern Florida</article-title><source>Viruses</source><volume>12</volume><person-group person-group-type="author"><name><surname>Quaglia</surname><given-names>A.I.</given-names></name><name><surname>Blosser</surname><given-names>E.M.</given-names></name><name><surname>McGregor</surname><given-names>B.L.</given-names></name><name><surname>Runkel</surname><given-names>A.E.</given-names></name><name><surname>Sloyer</surname><given-names>K.E.</given-names></name><name><surname>Erram</surname><given-names>D.</given-names></name><name><surname>Wisely</surname><given-names>S.M.</given-names></name><name><surname>Burkett-Cadena</surname><given-names>N.D.</given-names></name></person-group><year>2020</year><fpage>1</fpage><lpage>25</lpage><page-range>1-25</page-range><pub-id pub-id-type="doi">10.3390/v12090931</pub-id></element-citation></ref><ref id="BIBR-57"><element-citation publication-type="journal"><article-title>R: a language and environment for statistical computing</article-title><person-group person-group-type="author"><name><surname>Team</surname><given-names>R.Core</given-names></name></person-group><year>2025</year><comment>Available at:</comment><ext-link xlink:href="https://www.r-project.org/" ext-link-type="uri">https://www.r-project.org/</ext-link></element-citation></ref><ref id="BIBR-58"><element-citation publication-type="article-journal"><article-title>Rainforest conversion to rubber and oil palm reduces abundance, biomass and diversity of canopy spiders</article-title><source>PeerJ</source><volume>1</volume><person-group person-group-type="author"><name><surname>Ramos</surname><given-names>D.</given-names></name><name><surname>Hartke</surname><given-names>T.R.</given-names></name><name><surname>Buchori</surname><given-names>D.</given-names></name><name><surname>Dupérré</surname><given-names>N.</given-names></name><name><surname>Hidayat</surname><given-names>P.</given-names></name><name><surname>Lia</surname><given-names>M.</given-names></name><name><surname>Harms</surname><given-names>D.</given-names></name><name><surname>Scheu</surname><given-names>S.</given-names></name><name><surname>Drescher</surname><given-names>J.</given-names></name></person-group><year>2022</year><fpage>1</fpage><lpage>27</lpage><page-range>1-27</page-range><pub-id pub-id-type="doi">10.7717/peerj.13898</pub-id></element-citation></ref><ref id="BIBR-59"><element-citation publication-type="article-journal"><article-title>Plant diversity, forest dependency, and alien plant invasions in tropical agricultural landscapes</article-title><source>Biological Conservation</source><volume>213</volume><person-group person-group-type="author"><name><surname>Rembold</surname><given-names>K.</given-names></name><name><surname>Mangopo</surname><given-names>H.</given-names></name><name><surname>Tjitrosoedirdjo</surname><given-names>S.S.</given-names></name><name><surname>Kreft</surname><given-names>H.</given-names></name></person-group><year>2017</year><fpage>234</fpage><lpage>242</lpage><page-range>234-242</page-range><pub-id pub-id-type="doi">10.1016/j.biocon.2017.07.020</pub-id></element-citation></ref><ref id="BIBR-60"><element-citation publication-type="article-journal"><article-title>Effects of tree and herb biodiversity on Diptera, a hyperdiverse insect order</article-title><source>Oecologia</source><volume>174</volume><person-group person-group-type="author"><name><surname>Scherber</surname><given-names>C.</given-names></name><name><surname>Vockenhuber</surname><given-names>E.A.</given-names></name><name><surname>Stark</surname><given-names>A.</given-names></name><name><surname>Meyer</surname><given-names>H.</given-names></name><name><surname>Tscharntke</surname><given-names>T.</given-names></name></person-group><year>2014</year><pub-id pub-id-type="doi">10.1007/s00442-013-2865-7</pub-id></element-citation></ref><ref id="BIBR-61"><element-citation publication-type="book"><article-title>Insect Ecology, An Ecosystem Approach</article-title><person-group person-group-type="author"><name><surname>Schowalter</surname><given-names>D.</given-names></name></person-group><year>2011</year><page-range>978-0-12-381351-0</page-range><publisher-name>Academic Press. ISBN</publisher-name><publisher-loc>San Diego</publisher-loc></element-citation></ref><ref id="BIBR-62"><element-citation publication-type="article-journal"><article-title>Diptera species and functional diversity across tropical Australian countryside landscapes</article-title><source>Biological Conservation</source><volume>191</volume><person-group person-group-type="author"><name><surname>Smith</surname><given-names>T.J.</given-names></name><name><surname>Mayfield</surname><given-names>M.M.</given-names></name></person-group><year>2015</year><fpage>436</fpage><lpage>443</lpage><page-range>436-443</page-range><pub-id pub-id-type="doi">10.1016/j.biocon.2015.07.035</pub-id></element-citation></ref><ref id="BIBR-63"><element-citation publication-type="article-journal"><article-title>Abundance, distribution mapping and DNA barcoding of Procontarinia robusta (Diptera: Cecidomyiidae), a mango gall midge in Bali, Indonesia</article-title><source>Biodiversitas Journal of Biological Diversity</source><volume>23</volume><person-group person-group-type="author"><name><surname>Susila</surname><given-names>I.W.</given-names></name><name><surname>Sumiartha</surname><given-names>I.K.</given-names></name><name><surname>Supartha</surname><given-names>I.W.</given-names></name><name><surname>Yudha</surname><given-names>I.K.W.</given-names></name><name><surname>Utama</surname><given-names>I.W.E.K.</given-names></name><name><surname>Yasa</surname><given-names>I.W.S.</given-names></name><name><surname>Wiradana</surname><given-names>P.A.</given-names></name></person-group><year>2022</year><fpage>6428</fpage><lpage>6436</lpage><page-range>6428-6436</page-range><pub-id pub-id-type="doi">10.13057/biodiv/d231241</pub-id></element-citation></ref><ref id="BIBR-64"><element-citation publication-type="article-journal"><article-title>Contribution of epiphytes on the canopy insect population in oil palm plantations in North Sumatera</article-title><source>ARPN Journal of Engineering and Applied Sciences</source><volume>11</volume><person-group person-group-type="author"><name><surname>Suzanti</surname><given-names>F.</given-names></name><name><surname>Kuswardani</surname><given-names>R.A.</given-names></name><name><surname>Rahayu</surname><given-names>S.</given-names></name><name><surname>Susanto</surname><given-names>A.</given-names></name></person-group><year>2016</year><fpage>6982</fpage><lpage>6998</lpage><page-range>6982-6998</page-range></element-citation></ref><ref id="BIBR-65"><element-citation publication-type="article-journal"><article-title>Karakteristik lingkungan terhadap komunitas serangga</article-title><source>Journal of Natural Resources and Environmental Management</source><volume>9</volume><person-group person-group-type="author"><name><surname>Taradipha</surname><given-names>M.R.R.</given-names></name><name><surname>Rushayati</surname><given-names>S.B.</given-names></name><name><surname>Haneda</surname><given-names>N.F.</given-names></name></person-group><year>2018</year><fpage>394</fpage><lpage>404</lpage><page-range>394-404</page-range><pub-id pub-id-type="doi">10.29244/jpsl.9.2.394-404</pub-id></element-citation></ref><ref id="BIBR-66"><element-citation publication-type="article-journal"><article-title>Experimental biodiversity enrichment in oil-palm-dominated landscapes in Indonesia</article-title><source>Frontiers in Plant Science</source><volume>7</volume><person-group person-group-type="author"><name><surname>Teuscher</surname><given-names>M.</given-names></name><name><surname>Gerard</surname><given-names>A.</given-names></name><name><surname>Brose</surname><given-names>U.</given-names></name><name><surname>Buchori</surname><given-names>D.</given-names></name><name><surname>Clough</surname><given-names>Y.</given-names></name><name><surname>Ehbrecht</surname><given-names>M.</given-names></name><name><surname>Holscher</surname><given-names>D.</given-names></name><name><surname>Irawan</surname><given-names>B.</given-names></name><name><surname>Sundawati</surname><given-names>L.</given-names></name><name><surname>Wollni</surname><given-names>M.</given-names></name><name><surname>Kreft</surname><given-names>H.</given-names></name></person-group><year>2016</year><fpage>1</fpage><lpage>15</lpage><page-range>1-15</page-range><pub-id pub-id-type="doi">10.3389/fpls.2016.01538</pub-id></element-citation></ref><ref id="BIBR-67"><element-citation publication-type="article-journal"><article-title>Landscape and farm-level management for conservation of potential pollinators in Indonesian cocoa agroforests</article-title><source>Biological Conservation</source><volume>257</volume><person-group person-group-type="author"><name><surname>Toledo-Hernandez</surname><given-names>M.</given-names></name><name><surname>Tscharntke</surname><given-names>T.</given-names></name><name><surname>Tjoa</surname><given-names>A.</given-names></name><name><surname>Anshary</surname><given-names>A.</given-names></name><name><surname>Cyio</surname><given-names>B.</given-names></name><name><surname>Wanger</surname><given-names>T.C.</given-names></name></person-group><year>2021</year><fpage>1</fpage><lpage>8</lpage><page-range>1-8</page-range><pub-id pub-id-type="doi">10.1016/j.biocon.2021.109106</pub-id></element-citation></ref><ref id="BIBR-68"><element-citation publication-type="article-journal"><article-title>Deforestation homogenizes tropical parasitoid-host networks</article-title><source>Ecology</source><volume>91</volume><person-group person-group-type="author"><name><surname>Laliberté</surname><given-names>E.</given-names></name><name><surname>Tylianakis</surname><given-names>J.M.</given-names></name></person-group><year>2010</year><fpage>1740</fpage><lpage>1747</lpage><page-range>1740-1747</page-range><pub-id pub-id-type="doi">10.1890/09-1328.1</pub-id></element-citation></ref><ref id="BIBR-69"><element-citation publication-type="journal"><article-title>Ggplot2: elegant graphics for data analysis</article-title><person-group person-group-type="author"><name><surname>Wickham</surname><given-names>H.</given-names></name></person-group><year>2016</year><publisher-name>Springer-Verlag</publisher-name><publisher-loc>New York</publisher-loc><comment>DOI:</comment><pub-id pub-id-type="doi">10.1007/978-3-319-24277-4_9</pub-id></element-citation></ref><ref id="BIBR-70"><element-citation publication-type="article-journal"><article-title>Field observations on Ceratopogonidae and other Diptera: Nematocera associated with cocoa flowers in Brazil</article-title><source>Bulletin of Entomological Research</source><volume>67</volume><person-group person-group-type="author"><name><surname>Winder</surname><given-names>J.A.</given-names></name></person-group><year>1977</year><fpage>57</fpage><lpage>63</lpage><page-range>57-63</page-range><pub-id pub-id-type="doi">10.1017/S0007485300010890</pub-id></element-citation></ref><ref id="BIBR-71"><element-citation publication-type="article-journal"><article-title>Mixed-species tree plantings enhance structural complexity in oil palm plantations</article-title><source>Agriculture, Ecosystems and Environment</source><volume>283</volume><person-group person-group-type="author"><name><surname>Zemp</surname><given-names>D.C.</given-names></name><name><surname>Ehbrecht</surname><given-names>M.</given-names></name><name><surname>Seidel</surname><given-names>D.</given-names></name><name><surname>Ammer</surname><given-names>C.</given-names></name><name><surname>Craven</surname><given-names>D.</given-names></name><name><surname>Erkelenz</surname><given-names>J.</given-names></name><name><surname>Irawan</surname><given-names>B.</given-names></name><name><surname>Sundawati</surname><given-names>L.</given-names></name><name><surname>Hölscher</surname><given-names>D.</given-names></name><name><surname>Kreft</surname><given-names>H.</given-names></name></person-group><year>2019</year><fpage>1</fpage><lpage>9</lpage><page-range>1-9</page-range><pub-id pub-id-type="doi">10.1016/j.agee.2019.06.003</pub-id></element-citation></ref><ref id="BIBR-72"><element-citation publication-type="article-journal"><article-title>Tree islands enhance biodiversity and functioning in oil palm landscapes</article-title><source>Nature</source><volume>618</volume><person-group person-group-type="author"><name><surname>Zemp</surname><given-names>D.C.</given-names></name><name><surname>Guerrero-Ramirez</surname><given-names>N.</given-names></name><name><surname>Brambach</surname><given-names>F.</given-names></name><name><surname>Darras</surname><given-names>K.</given-names></name><name><surname>Grass</surname><given-names>I.</given-names></name><name><surname>Potapov</surname><given-names>A.</given-names></name><name><surname>Röll</surname><given-names>A.</given-names></name><name><surname>Arimond</surname><given-names>I.</given-names></name><name><surname>Ballauff</surname><given-names>J.</given-names></name><name><surname>Behling</surname><given-names>H.</given-names></name><name><surname>Berkelmann</surname><given-names>D.</given-names></name><name><surname>Biagioni</surname><given-names>S.</given-names></name><name><surname>Buchori</surname><given-names>D.</given-names></name><name><surname>Craven</surname><given-names>D.</given-names></name><name><surname>Daniel</surname><given-names>R.</given-names></name><name><surname>Gailing</surname><given-names>O.</given-names></name><name><surname>Ellsäßer</surname><given-names>F.</given-names></name><name><surname>Fardiansah</surname><given-names>R.</given-names></name><name><surname>Hennings</surname><given-names>N.</given-names></name><name><surname>Irawan</surname><given-names>B.</given-names></name><name><surname>Khokthong</surname><given-names>W.</given-names></name><name><surname>Krashevska</surname><given-names>V.</given-names></name><name><surname>Krause</surname><given-names>A.</given-names></name><name><surname>Kückes</surname><given-names>J.</given-names></name><name><surname>Li</surname><given-names>K.</given-names></name><name><surname>Lorenz</surname><given-names>H.</given-names></name><name><surname>Maraun</surname><given-names>M.</given-names></name><name><surname>Merk</surname><given-names>M.S.</given-names></name><name><surname>Moura</surname><given-names>C.C.M.</given-names></name><name><surname>Mulyani</surname><given-names>Y.A.</given-names></name><name><surname>Paterno</surname><given-names>G.B.</given-names></name><name><surname>Pebrianti</surname><given-names>H.D.</given-names></name><name><surname>Polle</surname><given-names>A.</given-names></name><name><surname>Prameswari</surname><given-names>D.A.</given-names></name><name><surname>Sachsenmaier</surname><given-names>L.</given-names></name><name><surname>Scheu</surname><given-names>S.</given-names></name><name><surname>Schneider</surname><given-names>D.</given-names></name><name><surname>Setiajiati</surname><given-names>F.</given-names></name><name><surname>Setyaningsih</surname><given-names>C.A.</given-names></name><name><surname>Sundawati</surname><given-names>L.</given-names></name><name><surname>Tscharntke</surname><given-names>T.</given-names></name><name><surname>Wollni</surname><given-names>M.</given-names></name><name><surname>Hölscher</surname><given-names>D.</given-names></name><name><surname>Kreft</surname><given-names>H.</given-names></name></person-group><year>2023</year><fpage>316</fpage><lpage>321</lpage><page-range>316-321</page-range><pub-id pub-id-type="doi">10.1038/s41586-023-06086-5</pub-id></element-citation></ref><ref id="BIBR-73"><element-citation publication-type="article-journal"><article-title>Larval development sites of the main Culicoides species (Diptera: Ceratopogonidae) in northern Europe and distribution of coprophilic species larvae in Belgian pastures</article-title><source>Veterinary Parasitology</source><volume>205</volume><person-group person-group-type="author"><name><surname>Zimmer</surname><given-names>J.</given-names></name><name><surname>Brostaux</surname><given-names>Y.</given-names></name><name><surname>Haubruge</surname><given-names>E.</given-names></name><name><surname>Francis</surname><given-names>F.</given-names></name></person-group><year>2014</year><fpage>676</fpage><lpage>686</lpage><page-range>676-686</page-range><pub-id pub-id-type="doi">10.1016/j.vetpar.2014.08.029</pub-id></element-citation></ref></ref-list></back></article>