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<article xmlns:xlink="http://www.w3.org/1999/xlink" dtd-version="1.3" article-type="research-article"><front><journal-meta><journal-id journal-id-type="issn">2089-0257</journal-id><journal-title-group><journal-title>Jurnal Entomologi Indonesia</journal-title></journal-title-group><issn pub-type="epub">2089-0257</issn><issn pub-type="ppub">1829-7722</issn><publisher><publisher-name>Perhimpunan Entomologi Indonesia</publisher-name></publisher></journal-meta><article-meta><article-id pub-id-type="doi">10.5994/jei.20.1.49</article-id><article-categories><subj-group subj-group-type="toc-heading"><subject>PENDAHULUAN</subject></subj-group><subj-group subj-group-type="toc-heading"><subject>BAHAN DAN METODE</subject><subj-group subj-group-type="toc-heading"><subject>Waktu dan tempat penelitian</subject></subj-group><subj-group subj-group-type="toc-heading"><subject>Desain dan bahan penelitian</subject></subj-group><subj-group subj-group-type="toc-heading"><subject>Pemeliharaan lipas jerman</subject></subj-group><subj-group subj-group-type="toc-heading"><subject>Pengujian</subject></subj-group><subj-group subj-group-type="toc-heading"><subject>Pengamatan dan kriteria pengujian</subject></subj-group><subj-group subj-group-type="toc-heading"><subject>Analisis data</subject></subj-group></subj-group><subj-group subj-group-type="toc-heading"><subject>HASIL</subject></subj-group><subj-group subj-group-type="toc-heading"><subject>PEMBAHASAN</subject></subj-group><subj-group subj-group-type="toc-heading"><subject>KESIMPULAN</subject></subj-group></article-categories><title-group><article-title>Pengendalian lipas jerman (Blattella germanica Linnaeus) dengan menggunakan attractive toxic sugar baits</article-title><subtitle>Control of the german cockroach (Blattella germanica Linnaeus) by using attractive toxic sugar baits</subtitle></title-group><contrib-group><contrib contrib-type="author"><name><surname>Khairi</surname><given-names>Fahmi</given-names></name><address><country>Indonesia</country><email>upikke@apps.ipb.ac.id</email></address><xref ref-type="aff" rid="AFF-1"/></contrib><contrib contrib-type="author"><name><surname>Hadi</surname><given-names>Upik Kesumawati</given-names></name><address><country>Indonesia</country><email>upikke@apps.ipb.ac.id</email></address><xref ref-type="aff" rid="AFF-2"/></contrib><contrib contrib-type="author"><name><surname>Amin</surname><given-names>Akhmad Arif</given-names></name><address><country>Indonesia</country><email>aaamin@apps.ipb.ac.id</email></address><xref ref-type="aff" rid="AFF-2"/></contrib><aff id="AFF-1">Sekolah Pascasarjana, Peminatan Parasitologi dan Entomologi Kesehatan, IPB University, Indonesia</aff><aff id="AFF-2">Divisi Parasitologi dan Entomologi Kesehatan, Sekolah Kedokteran Hewan dan Biomedis,  IPB University, Indonesia</aff></contrib-group><pub-date date-type="pub" iso-8601-date="2023-7-11" publication-format="electronic"><day>11</day><month>7</month><year>2023</year></pub-date><volume>20</volume><issue>1</issue><fpage>49</fpage><history><date date-type="received" iso-8601-date="2022-12-7"><day>7</day><month>12</month><year>2022</year></date><date date-type="accepted" iso-8601-date="2023-7-7"><day>7</day><month>7</month><year>2023</year></date></history><permissions><copyright-statement>Copyright (c) 2023 Fahmi  Khairi, Upik Kesumawati Hadi, Akhmad Arif  Amin</copyright-statement><license license-type="open-access"><ali:license_ref xmlns:ali="http://www.niso.org/schemas/ali/1.0/">https://creativecommons.org/licenses/by/4.0</ali:license_ref><license-p>This work is licensed under a Creative Commons Attribution 4.0 International License.Authors who publish with this journal agree to the following terms:

Authors retain copyright and grant the journal right of first publication with the work simultaneously licensed under a Creative Commons Attribution 4.0 International License that allows others to share the work with an acknowledgement of the work's authorship and initial publication in this journal.
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Authors are permitted and encouraged to post their work online (e.g., in institutional repositories or on their website) prior to and during the submission process, as it can lead to productive exchanges, as well as earlier and greater citation of published work (See The Effect of Open Access).</license-p></license></permissions><self-uri xlink:href="https://jurnal.pei-pusat.org/index.php/jei/article/view/755">https://jurnal.pei-pusat.org/index.php/jei/article/view/755</self-uri><abstract><p>The use of attractive toxic sugar baits (ATSB) is a novelty in vector and residential pest control. This study aims to determine the concentration of sugar bait that is able to attract the german cockroach <italic>Blattela germanica </italic>Linnaeus, and to determine the effectiveness of various types of toxic substances as an ATSB to control german cockroaches. This method is based on the behavior of adult insects that eat sugars from sources that contain toxic materials and will ultimately kill the insects. The types of toxic materials used in this study were inorganic insecticides (boric acid), pyrrole (chlorefenapir), neonicotinoids (dinotefuran), and pyrethroids (permethrin and deltamethrin). The results showed that an attractive sugar bait for german cockroach was 5% sucrose. ATSB with 25% boric acid toxic material caused 46.25% of cockroach mortality, and 0.12% chlorfenapir caused 70% of cockroach mortality 24 hours after treatment. Furthermore, ATSB with dinetofuran 0.08% and deltamethrin in various concentrations caused the highest cockroach mortality (100%), while permethrin caused cockroach mortality which was lower than ATSB with other toxic materials at the same time of observation. The conclusions of this study provide basic information about the use of ATSB and its potential in controlling the german cockroach in Indonesia.</p></abstract><kwd-group><kwd>ATSB (called attractive toxic sugar baits)</kwd><kwd>Blattella germanica</kwd><kwd>cockroach</kwd><kwd>control</kwd></kwd-group></article-meta></front><body><sec><title>PENDAHULUAN</title><p>Lipas merupakan satu di antara serangga primitif yang termasuk ke dalam Ordo Dyctioptera. Lipas termasuk serangga omnivora dengan tipe mulut penggigit-pengunyah (biting-chewing mouthparts) yang memakan hampir semua zat organik yang ditanam, diproduksi, disimpan, dikeluarkan, atau dibuang oleh manusia <xref ref-type="bibr" rid="BIBR-11">(Hadi &amp; Soviana, 2010)</xref><xref ref-type="bibr" rid="BIBR-6">(Brenner &amp; Kramer, 2019)</xref>.</p><p>Keberadaan lipas jerman (Blattella germanica Linnaeus) di rumah dan dapur komersial dapat menjadi indikator sanitasi atau tata kelola rumah tangga yang buruk. Selain sebagai hama pengganggu, lipas juga memiliki peran penting dalam dunia kesehatan. Persediaan makanan dan peralatan berisiko terkontaminasi oleh patogen penyakit yang dapat ditransmisikan oleh lipas. Manusia juga dapat terpapar protein alergen dari lipas yang berpotensi menyebabkan alergi dan dapat menyebabkan penyakit pernapasan <xref ref-type="bibr" rid="BIBR-26">(Sheih et al., 2017)</xref><xref ref-type="bibr" rid="BIBR-6">(Brenner &amp; Kramer, 2019)</xref>.</p><p>Lipas jerman merupakan satu di antara hama permukiman yang paling penting. Lipas ini di- temukan di berbagai lokasi strategis di perkotaan. Lipas jerman dilaporkan ditemukan di hotel dan restoran <xref ref-type="bibr" rid="BIBR-17">(Lee et al., 1999)</xref>; kapal dan pesawat terbang <xref ref-type="bibr" rid="BIBR-29">(Song et al., 2003)</xref>; rumah sakit <xref ref-type="bibr" rid="BIBR-20">(Pai et al., 2010)</xref><xref ref-type="bibr" rid="BIBR-28">(Sinulingga, 2020)</xref>; asrama <xref ref-type="bibr" rid="BIBR-25">(Shahraki, 2013)</xref>; bus <xref ref-type="bibr" rid="BIBR-24">(Rafiuddin, 2016)</xref>; kapal barang dan kapal penumpang <xref ref-type="bibr" rid="BIBR-2">(Alias, 2018)</xref><xref ref-type="bibr" rid="BIBR-31">(Supriyanto, 2018)</xref>.</p><p>Umumnya, pengendalian lipas masih menggunakan insektisida yang mempunyai dampak negatif, seperti resistensi, yang dapat memengaruhi kesuksesan sebuah program pengendalian. Resistensi lipas jerman terhadap berbagai golongan insektisida (organofosfat, karbamat, fenilpirasol, dan piretroid) dilaporkan oleh beberapa peneliti, seperti <xref ref-type="bibr" rid="BIBR-20">(Pai et al., 2010)</xref>, <xref ref-type="bibr" rid="BIBR-23">(Rahayu, 2012)</xref>, dan <xref ref-type="bibr" rid="BIBR-15">(Khani et al., 2020)</xref>.</p><p>Penggunaan attractive toxic sugar baits (ATSB) merupakan hal baru dalam pengendalian vektor dan hama permukiman. Prinsip penggunaan ATSB didasarkan pada perilaku serangga yang membutuhkan gula agar memakan gula dari sumber yang mengandung bahan toksik yang dapat membunuh serangga tersebut <xref ref-type="bibr" rid="BIBR-16">(Kline et al., 2018)</xref>.</p><p>Laporan penggunaan ATSB dalam pengendalian vektor dan hama permukiman menunjukkan kesuksesan. Beberapa bukti menunjukkan keberhasilan dalam pengendalian nyamuk hingga berkurang secara signifikan, seperti yang dilaporkan oleh <xref ref-type="bibr" rid="BIBR-14">(Khallayoune et al., 2013)</xref>, <xref ref-type="bibr" rid="BIBR-21">(Qualls et al., 2014)</xref>, <xref ref-type="bibr" rid="BIBR-1">(Abdullah et al., 2019)</xref>. Penggunaan ATSB juga menunjukkan keberhasilan dalam pengendalian lalat rumah (Musca domestica Linneaus), seperti yang dilaporkan <xref ref-type="bibr" rid="BIBR-4">(Amin et al., 2017)</xref>. Dampak minimal penggunaan ATSB terhadap organisme nontarget juga telah dilaporkan oleh <xref ref-type="bibr" rid="BIBR-14">(Khallayoune et al., 2013)</xref> dan <xref ref-type="bibr" rid="BIBR-21">(Qualls et al., 2014)</xref> pada Ordo Hymenoptera, Lepidoptera, Coleoptera, Diptera, Hemiptera, Orthoptera, dan Neuroptera. Meskipun demikian, laporan pemanfaatan ATSB untuk mengendalikan populasi lipas belum ada.</p><p>Penelitian ini bertujuan untuk menentukan konsentrasi umpan gula yang mampu menarik lipas jerman <italic>B. germanica, </italic>dan menentukan keefektifan berbagai jenis bahan toksik sebagai ATSB dalam pengendalian lipas jerman.</p></sec><sec><title>BAHAN DAN METODE</title><sec><title>Waktu dan tempat penelitian</title><p>Penelitian dilaksanakan pada bulan September 2021 hingga Maret 2022 di Laboratorium Entomologi Kesehatan, Departemen Ilmu Penyakit Hewan dan Kesehatan Masyarakat Veteriner, Sekolah Kedokteran Hewan dan Biomedis, IPB University.</p></sec><sec><title>Desain dan bahan penelitian</title><p>Penelitian eksperimental ini dilakukan di laboratorium dengan menggunakan rancangan acak lengkap. Penelitian diawali dengan pemeliharaan lipas jerman <italic>strain </italic>rentan yang berasal dari Laboratorium Entomologi Kesehatan, Sekolah Kedokteran Hewan Biomedis (SKHB), IPB University. Lipas hasil pemeliharaan kemudian diujikan pada berbagai jenis ATSB. Gula sebagai atraktan yang digunakan dalam penelitian ini adalah sukrosa (GULAKU®). Adapun bahan toksik yang digunakan dalam penelitian ini adalah asam borat, klorfenapir (MYTHIC 240 SC®), dinotefuran (SECLIRA 40 SG®), permetrin (KLENSECT 200 EC®), dan deltametrin (DELTAMETRIN 98 TC®).</p></sec><sec><title>Pemeliharaan lipas jerman</title><p>Lipas jerman dipelihara di Insektarium Laboratorium Entomologi Kesehatan, SKHB IPB University. Lipas dipelihara dalam wadah plastik (volume ± 12 l). Pemeliharaan lipas dilakukan di ruangan dengan suhu kamar (26–30 °C), kelembaban 70–90% dan diberi pakan berupa pellet pakan ikan serta disediakan air minum ad libitum. Pemeliharaan lipas dimulai dari tahapan telur (ooteka) sampai menjadi dewasa.</p></sec><sec><title>Pengujian</title><p>Lipas jerman yang digunakan pada pengujian adalah lipas jantan dan betina dewasa umur 3–5 hari hasil pemeliharaan. Sebelum pengujian dimulai, lipas yang akan diujikan dipuasakan selama 24 jam dan dilakukan aklimatisasi selama 1 jam dalam kotak pengujian (30 cm x 30 cm x 30 cm) untuk memastikan bahwa kondisi sama dan tidak berpengaruh terhadap kematian lipas uji. Pengujian pertama adalah uji ketertarikan yang dilakukan untuk mengetahui konsentrasi umpan gula yang dapat menarik lipas jerman. Umpan gula berbagai konsentrasi (1,25%, 2,5%, 5%, dan 10%) dan kontrol (tanpa gula) disiapkan dan diletakkan dalam cawan petri. Kemudian sebanyak 20 lipas jerman dewasa (jantan dan betina) dimasukkan ke dalam kotak pengujian. Larutan umpan gula dimasukkan ke dalam kotak pengujian sebanyak 20 ml dan dibiarkan selama 1 jam. Konsentrasi umpan gula yang menarik bagi lipas kemudian digunakan untuk pengujian selanjutnya.</p><p>Pengujian kedua adalah uji keefektifan ATSB dengan mencampurkan umpan gula dengan beberapa bahan toksik berbagai konsentrasi. Perlakuan pertama mencampurkan umpan gula dengan bahan toksik asam borat konsentrasi 0,25%, 0,5%, 1%, 2,5%, dan 5%. Perlakuan kedua mencampurkan umpan gula dengan bahan toksik klorfenapir konsentrasi 0,12%, 0,24%, dan 0,36%. Perlakuan ketiga mencampurkan umpan gula dengan bahan toksik dinotefuran konsentrasi 0,08%, 0,16%, dan 0,32%. Perlakuan keempat mencampurkan umpan gula dengan bahan toksik permetrin konsentrasi 0,01%, 0,02%, dan 0,04%. Perlakuan kelima mencampurkan umpan gula dengan bahan toksik deltametrin konsentrasi 1,96%, 3,92%, dan 7,84%. Setiap perlakuan ditambahkan kontrol berupa larutan umpan gula 5% tanpa bahan toksik. Setiap konsentrasi berbagai bahan toksik dilakukan ulangan sebanyak empat kali. Sebanyak 20 lipas jerman dewasa (jantan dan betina) dimasukkan ke dalam kotak pengujian. Larutan ATSB sebanyak 20 ml dimasukkan ke dalam kotak pengujian dan dibiarkan selama 1 jam.</p></sec><sec><title>Pengamatan dan kriteria pengujian</title><p>Pengamatan uji ketertarikan terhadap umpan gula dilakukan dengan menghitung lipas yang datang mendekati umpan pada menit ke-15, 30, 45, dan 60 selama 2 menit<xref ref-type="bibr" rid="BIBR-5">(Az et al., 2018)</xref>. Data kedatangan lipas dihitung secara kumulatif dalam satu jam dan dirata-ratakan. Adapun pengamatan terhadap uji keefektifan ATSB dilakukan dengan melihat kematian lipas pada jam ke-1, 6, 24, dan 48 pascaperlakuan.</p><p>Kriteria uji ketertarikan terhadap umpan gula ditentukan berdasarkan jumlah kedatangan lipas terbanyak selama 1 jam pengamatan <xref ref-type="bibr" rid="BIBR-5">(Az et al., 2018)</xref> Kriteria uji keefektifan ATSB yang diuji ditentukan berdasarkan persentase kematian lipas dalam waktu 24 jam. Kriteria ATSB dikatakan efektif apabila angka kematian lipas berkisar antara 90–100%.</p></sec><sec><title>Analisis data</title><p>Data penelitian ditabulasi dengan menggunakan aplikasi Microsoft Excel 2016 dan Minitab® 18 version. Selanjutnya data hasil penelitian dianalisis secara statistik menggunakan analisis ragam ANOVA.</p></sec></sec><sec><title>HASIL</title><p>Berdasarkan hasil penelitian, konsentrasi umpan gula 5% paling menarik bagi lipas jerman. Total kedatangan lipas jerman pada konsentrasi 5% sebanyak 66 kedatangan dalam waktu 1 jam pengamatan (rata-rata 16,5 ± 1,00 lipas). Nilai ini berbeda nyata dibandingkan dengan kontrol dan konsentrasi umpan gula lainnya<xref ref-type="table" rid="table-3bbdad7a">Table 1.</xref>.</p><p>Hasil pengujian efektivitas ATSB dengan mencampurkan bahan toksik asam borat konsentrasi 0,25% menunjukkan angka kematian lipas jerman sebesar 46,25% pada 24 jam pascaperlakuan. Pengujian ATSB berupa umpan gula dengan bahan toksik klorfenapir 0,12% memperlihatkan angka kematian lipas jerman sebesar 70% pada 24 jam pascaperlakuan. Adapun angka kematian lipas jerman mencapai 100% dalam waktu 24 jam pascaperlakuan ATSB berupa umpan gula dengan bahan toksik dinotefuran 0,08%. Pengujian ATSB berupa umpan gula dengan bahan toksik permetrin menunjukkan angka kematian yang lebih rendah (22,5–52,5%) dibandingkan dengan bahan toksik lainnya. Angka kematian lipas jerman sebesar 52,5% pada permetrin dengan konsentrasi 0,04% dalam waktu 24 jam pascaperlakuan. Sementara itu, angka kematian lipas jerman mencapai 100% dalam waktu 24 jam pascaperlakuan ATSB berupa umpan gula dengan bahan toksik deltametrin berbagai konsentrasi (1,96%, 3,92%, 7,84%)<xref ref-type="table" rid="table-57ddce80">Tabel 2.</xref>.</p><table-wrap id="table-3bbdad7a"><label>Table 1.</label><caption><p>Angka kedatangan lipas Blattella germanica pada pengujianketertarikan terhadap umpan gula<italic>(Number of arrival Blattella germanica in attractiveness test for sugar baits)</italic></p></caption><table frame="box" rules="all"><thead><tr><th colspan="1" rowspan="1" style="" align="left" valign="middle"><p>Konsentrasi umpan gula <italic>(Concentration of sugar bait)</italic> (%)</p></th><th colspan="1" rowspan="1" style="" align="center" valign="middle"><p>Total kedatangan lipas dari empat ulangan (dalam 1 jam)</p><p><italic>(Total arrival of cockroaches from four replicates (in 1 hour))</italic></p></th><th colspan="1" rowspan="1" style="" align="center" valign="middle"><p>Rata-rata kedatangan lipas* (dalam 1 jam)</p><p><italic>(Average cockroach arrival* (in 1 hour))</italic></p></th></tr></thead><tbody><tr><td colspan="1" rowspan="1" style="" align="left" valign="top"><p>0 (Kontrol)</p></td><td colspan="1" rowspan="1" style="" align="center" valign="top">2</td><td colspan="1" rowspan="1" style="" align="center" valign="top">0,5 ± 0,557 c</td></tr><tr><td colspan="1" rowspan="1" style="" align="left" valign="top"><p>1,25</p></td><td colspan="1" rowspan="1" style="" align="center" valign="top">22</td><td colspan="1" rowspan="1" style="" align="center" valign="top">5,5 ± 1,291 b</td></tr><tr><td colspan="1" rowspan="1" style="" align="left" valign="top"><p>2,5</p></td><td colspan="1" rowspan="1" style="" align="center" valign="top">30</td><td colspan="1" rowspan="1" style="" align="center" valign="top"><p>7,5 ± 4,43 b</p></td></tr><tr><td colspan="1" rowspan="1" style="" align="left" valign="top"><p>5</p></td><td colspan="1" rowspan="1" style="" align="center" valign="top">66</td><td colspan="1" rowspan="1" style="" align="center" valign="top"><p>16,5 ± 1,00 a</p></td></tr><tr><td colspan="1" rowspan="1" style="" align="left" valign="top"><p>10</p></td><td colspan="1" rowspan="1" style="" align="center" valign="top">29</td><td colspan="1" rowspan="1" style="" align="center" valign="top"><p>7,25 ± 1,258 b</p></td></tr></tbody></table><table-wrap-foot><p>*Angka yang diikuti oleh huruf yang sama pada kolom yang sama menunjukkan tidak berbeda nyata menurut uji Tukey (α = 0,05). <italic>(*Numbers followed by the same letter in the same column show no significant difference according to the Tukey test (α = 0,05)).</italic></p></table-wrap-foot></table-wrap><table-wrap id="table-57ddce80"><label>Tabel 2.</label><caption><p>Hasil pengujian kefektifan ATSB berbagai jenis bahan toksik terhadap lipas Blattella germanica<italic>(Results of testing the effectiveness of ATSB various types of toxic substances against the Blattella germanica cockroach)</italic></p></caption><table frame="box" rules="all"><thead><tr><th colspan="1" rowspan="2" style="" align="left" valign="middle"><p>Perlakuan <italic>(Treatment)</italic></p></th><th colspan="1" rowspan="2" style="" align="center" valign="middle"><p>Konsentrasi bahan toksik <italic>(Toxic concentration)</italic> (%)</p></th><th colspan="4" rowspan="1" style="" align="center" valign="middle"><p>Persentase kematian <italic>(Percentage of mortality)</italic></p><p>(%)</p></th></tr><tr><th colspan="1" rowspan="1" style="" align="center" valign="top"><p>1 jam</p><p><italic>(hour)</italic></p></th><th colspan="1" rowspan="1" style="" align="center" valign="top"><p>6 jam</p><p><italic>(hour)</italic></p></th><th colspan="1" rowspan="1" style="" align="center" valign="top"><p>24 jam</p><p><italic>(hour)</italic></p></th><th colspan="1" rowspan="1" style="" align="center" valign="top"><p>48 jam</p><p><italic>(hour)</italic></p></th></tr></thead><tbody><tr><td colspan="1" rowspan="1" style="" align="left" valign="top"><p>Kontrol: Gula 5% tanpa bahan toksik <italic>(Control: 5% sugar without toxic substances)</italic></p></td><td colspan="1" rowspan="1" style="" align="center" valign="top"><p>0</p></td><td colspan="1" rowspan="1" style="" align="center" valign="top">0</td><td colspan="1" rowspan="1" style="" align="center" valign="top">0</td><td colspan="1" rowspan="1" style="" align="center" valign="top">0</td><td colspan="1" rowspan="1" style="" align="center" valign="top">0</td></tr><tr><td colspan="1" rowspan="5" style="" align="left" valign="top"><p>Umpan gula 5% + Asam borat</p><p><italic>(Sugar baits 5% + Boric acid)</italic></p></td><td colspan="1" rowspan="1" style="" align="center" valign="top"><p>0,25</p></td><td colspan="1" rowspan="1" style="" align="center" valign="top">6,25</td><td colspan="1" rowspan="1" style="" align="center" valign="top">46,25</td><td colspan="1" rowspan="1" style="" align="center" valign="top">46,25</td><td colspan="1" rowspan="1" style="" align="center" valign="top"><p>90</p></td></tr><tr><td colspan="1" rowspan="1" style="" align="center" valign="top"><p>0,50</p></td><td colspan="1" rowspan="1" style="" align="center" valign="top">3,75</td><td colspan="1" rowspan="1" style="" align="center" valign="top"><p>7,5</p></td><td colspan="1" rowspan="1" style="" align="center" valign="top"><p>12,5</p></td><td colspan="1" rowspan="1" style="" align="center" valign="top"><p>77,5</p></td></tr><tr><td colspan="1" rowspan="1" style="" align="center" valign="top"><p>1,00</p></td><td colspan="1" rowspan="1" style="" align="center" valign="top">0,00</td><td colspan="1" rowspan="1" style="" align="center" valign="top"><p>2,5</p></td><td colspan="1" rowspan="1" style="" align="center" valign="top">11,25</td><td colspan="1" rowspan="1" style="" align="center" valign="top">46,25</td></tr><tr><td colspan="1" rowspan="1" style="" align="center" valign="top"><p>2,50</p></td><td colspan="1" rowspan="1" style="" align="center" valign="top">3,75</td><td colspan="1" rowspan="1" style="" align="center" valign="top"><p>5</p></td><td colspan="1" rowspan="1" style="" align="center" valign="top"><p>7,5</p></td><td colspan="1" rowspan="1" style="" align="center" valign="top">41,25</td></tr><tr><td colspan="1" rowspan="1" style="" align="center" valign="top"><p>5,00</p></td><td colspan="1" rowspan="1" style="" align="center" valign="top">2,50</td><td colspan="1" rowspan="1" style="" align="center" valign="top">6,25</td><td colspan="1" rowspan="1" style="" align="center" valign="top">8,75</td><td colspan="1" rowspan="1" style="" align="center" valign="top"><p>42,5</p></td></tr><tr><td colspan="1" rowspan="3" style="" align="left" valign="top"><p>Umpan gula 5% + Klorfenapir</p><p><italic>(Sugar baits 5% + Chlorfenapir)</italic></p></td><td colspan="1" rowspan="1" style="" align="center" valign="top"><p>0,12</p></td><td colspan="1" rowspan="1" style="" align="center" valign="top">5,00</td><td colspan="1" rowspan="1" style="" align="center" valign="top">21,25</td><td colspan="1" rowspan="1" style="" align="center" valign="top"><p>70</p></td><td colspan="1" rowspan="1" style="" align="center" valign="top">86,25</td></tr><tr><td colspan="1" rowspan="1" style="" align="center" valign="top"><p>0,24</p></td><td colspan="1" rowspan="1" style="" align="center" valign="top">1,25</td><td colspan="1" rowspan="1" style="" align="center" valign="top"><p>30</p></td><td colspan="1" rowspan="1" style="" align="center" valign="top">68,75</td><td colspan="1" rowspan="1" style="" align="center" valign="top"><p>85</p></td></tr><tr><td colspan="1" rowspan="1" style="" align="center" valign="top"><p>0,36</p></td><td colspan="1" rowspan="1" style="" align="center" valign="top">1,25</td><td colspan="1" rowspan="1" style="" align="center" valign="top"><p>22,5</p></td><td colspan="1" rowspan="1" style="" align="center" valign="top"><p>62,5</p></td><td colspan="1" rowspan="1" style="" align="center" valign="top">76,25</td></tr><tr><td colspan="1" rowspan="3" style="" align="left" valign="top"><p>Umpan gula 5% + Dinotefuran</p><p><italic>(Sugar 5% + Dinotefuran)</italic></p></td><td colspan="1" rowspan="1" style="" align="center" valign="top"><p>0,08</p></td><td colspan="1" rowspan="1" style="" align="center" valign="top"><p>70,00</p></td><td colspan="1" rowspan="1" style="" align="center" valign="top"><p>100</p></td><td colspan="1" rowspan="1" style="" align="center" valign="top"><p>100</p></td><td colspan="1" rowspan="1" style="" align="center" valign="top"><p>100</p></td></tr><tr><td colspan="1" rowspan="1" style="" align="center" valign="top"><p>0,16</p></td><td colspan="1" rowspan="1" style="" align="center" valign="top"><p>70,00</p></td><td colspan="1" rowspan="1" style="" align="center" valign="top"><p>100</p></td><td colspan="1" rowspan="1" style="" align="center" valign="top"><p>100</p></td><td colspan="1" rowspan="1" style="" align="center" valign="top"><p>100</p></td></tr><tr><td colspan="1" rowspan="1" style="" align="center" valign="top"><p>0,32</p></td><td colspan="1" rowspan="1" style="" align="center" valign="top"><p>60,0</p></td><td colspan="1" rowspan="1" style="" align="center" valign="top"><p>97,5</p></td><td colspan="1" rowspan="1" style="" align="center" valign="top"><p>100</p></td><td colspan="1" rowspan="1" style="" align="center" valign="top"><p>100</p></td></tr><tr><td colspan="1" rowspan="3" style="" align="left" valign="top"><p>Umpan gula 5% + Permetrin</p><p><italic>(Sugar baits 5% + Permethrin)</italic></p></td><td colspan="1" rowspan="1" style="" align="center" valign="top"><p>0,01</p></td><td colspan="1" rowspan="1" style="" align="center" valign="top">6,25</td><td colspan="1" rowspan="1" style="" align="center" valign="top"><p>22,5</p></td><td colspan="1" rowspan="1" style="" align="center" valign="top"><p>22,5</p></td><td colspan="1" rowspan="1" style="" align="center" valign="top"><p>25</p></td></tr><tr><td colspan="1" rowspan="1" style="" align="center" valign="top"><p>0,02</p></td><td colspan="1" rowspan="1" style="" align="center" valign="top">2,50</td><td colspan="1" rowspan="1" style="" align="center" valign="top"><p>42,5</p></td><td colspan="1" rowspan="1" style="" align="center" valign="top">43,75</td><td colspan="1" rowspan="1" style="" align="center" valign="top"><p>50</p></td></tr><tr><td colspan="1" rowspan="1" style="" align="center" valign="top"><p>0,04</p></td><td colspan="1" rowspan="1" style="" align="center" valign="top"><p>11,25</p></td><td colspan="1" rowspan="1" style="" align="center" valign="top">51,25</td><td colspan="1" rowspan="1" style="" align="center" valign="top"><p>52,5</p></td><td colspan="1" rowspan="1" style="" align="center" valign="top"><p>55</p></td></tr><tr><td colspan="1" rowspan="3" style="" align="left" valign="top"><p>Umpan gula 5% + Deltametrin</p><p><italic>(Sugar baits 5% + Deltamethrin)</italic></p></td><td colspan="1" rowspan="1" style="" align="center" valign="top"><p>1,96</p></td><td colspan="1" rowspan="1" style="" align="center" valign="top"><p>73,75</p></td><td colspan="1" rowspan="1" style="" align="center" valign="top"><p>100</p></td><td colspan="1" rowspan="1" style="" align="center" valign="top"><p>100</p></td><td colspan="1" rowspan="1" style="" align="center" valign="top"><p>100</p></td></tr><tr><td colspan="1" rowspan="1" style="" align="center" valign="top"><p>3,92</p></td><td colspan="1" rowspan="1" style="" align="center" valign="top"><p>81,25</p></td><td colspan="1" rowspan="1" style="" align="center" valign="top"><p>100</p></td><td colspan="1" rowspan="1" style="" align="center" valign="top"><p>100</p></td><td colspan="1" rowspan="1" style="" align="center" valign="top"><p>100</p></td></tr><tr><td colspan="1" rowspan="1" style="" align="center" valign="top"><p>7,84</p></td><td colspan="1" rowspan="1" style="" align="center" valign="top"><p>85</p></td><td colspan="1" rowspan="1" style="" align="center" valign="top"><p>100</p></td><td colspan="1" rowspan="1" style="" align="center" valign="top"><p>100</p></td><td colspan="1" rowspan="1" style="" align="center" valign="top"><p>100</p></td></tr></tbody></table></table-wrap></sec><sec><title>PEMBAHASAN</title><p>Karbohidrat merupakan komponen penting dalam tubuh serangga. Tubuh serangga dilapisi khitin yang termasuk ke dalam polisakarida. Karbohidrat adalah sumber energi utama dan komponen penting dari makanan sebagian besar serangga <xref ref-type="bibr" rid="BIBR-7">(Chapman, 1998)</xref><xref ref-type="bibr" rid="BIBR-16">(Kline et al., 2018)</xref>. Beberapa jenis karbohidrat, seperti glukosa, frukosa, sukrosa, dan karbohidrat jenis lainnya. digunakan sebagai umpan dalam beberapa penelitian. Adapun jenis karbohidrat yang digunakan dalam penelitian ini termasuk disakarida, yaitu sukrosa.</p><p>Konsentrasi sukrosa yang digunakan sebagai umpan dalam pengujian ATSB berkisar 5–20% <xref ref-type="bibr" rid="BIBR-8">(Fiorenzano et al., 2017)</xref>. Penggunaan sukrosa sebagai umpan gula dalam pengendalian hama permukiman dilaporkan oleh <xref ref-type="bibr" rid="BIBR-9">(Gore &amp; Schal, 2004)</xref> pada lipas jerman dengan konsentrasi 0,5% dan <xref ref-type="bibr" rid="BIBR-10">(Gu et al., 2020)</xref> dengan konsentrasi 8%. Penggunaan sukrosa 10% sebagai umpan gula dilaporkan oleh <xref ref-type="bibr" rid="BIBR-3">(Allan, 2011)</xref>, <xref ref-type="bibr" rid="BIBR-27">(Shin et al., 2011)</xref>, <xref ref-type="bibr" rid="BIBR-14">(Khallayoune et al., 2013)</xref>, <xref ref-type="bibr" rid="BIBR-4">(Amin et al., 2017)</xref>, dan <xref ref-type="bibr" rid="BIBR-18">(Maia et al., 2018)</xref>. Dalam penelitian ini konsentrasi gula yang digunakan untuk bahan pencampur dengan bahan toksik menjadi formula ATSB adalah konsentrasi umpan gula 5%. Konsentrasi ini yang paling menarik bagi lipas jerman untuk datang (66 lipas dalam waktu 1 jam pengamatan). Berdasarkan hasil penelitian tahap pertama ini, selanjutnya gula dengan konsentrasi 5% dipilih menjadi bahan formula ATSB yang selanjutnya ditambahkan dengan berbagai bahan toksik dari berbagai golongan insektisida mulai dari anorganik (seperti asam borat), pirol (klorfenafir), neonikotinoid (dinetofuran), dan piretroid (permetrin dan deltametrin).</p><p>Asam borat atau yang biasa dikenal asam boraks (H<sub>3</sub>BO<sub>3</sub>) merupakan senyawa yang biasa digunakan sebagai anti jamur dan anti bakteri di dunia medis. Asam borat termasuk insektisida anorganik yang cukup banyak digunakan pada tahun 1930-an dan kembali populer digunakan di masa sekarang. Umumnya, asam borat digunakan sebagai racun perut karena bersifat nonrepelen, dengan toksitas terhadap mamalia rendah, sangat toksik terhadap serangga, tidak berbau, tidak terserap kulit, dan <italic>slow acting</italic> <xref ref-type="bibr" rid="BIBR-32">(Wirawan, 2006)</xref><xref ref-type="bibr" rid="BIBR-12">(Committee, 2021)</xref>. Penggunaan asam borat dalam aplikasi ATSB dilaporkan oleh <xref ref-type="bibr" rid="BIBR-33">(Xue et al., 2006)</xref> pada berbagai jenis nyamuk. <xref ref-type="bibr" rid="BIBR-9">(Gore &amp; Schal, 2004)</xref> dan <xref ref-type="bibr" rid="BIBR-13">(Jiang et al., 2020)</xref> melaporkan penggunaan asam borat pada lipas jerman dengan konsentrasi asam borat yang tinggi dapat menyebabkan efek repelen. Hal ini menyebabkan perubahan rasa pada umpan sehingga menyebabkan terjadi penurunan konsumsi umpan oleh lipas jerman.</p><p>Klorfenapir merupakan insektisida dari golongan pirol yang bekerja dengan cara memisahkan rantai fosforilasi oksidatif di dalam mitokondria sehingga mengacaukan produksi <italic>adenosine triphosphate</italic> (ATP) pada serangga. Klorfenapir merupakan insektisida yang tidak mudah menguap, bersifat nonrepelen, dan toksisitas rendah terhadap mamalia <xref ref-type="bibr" rid="BIBR-32">(Wirawan, 2006)</xref><xref ref-type="bibr" rid="BIBR-12">(Committee, 2021)</xref>. Penggunaan klorfenapir dalam aplikasi ATSB dilaporkan <xref ref-type="bibr" rid="BIBR-3">(Allan, 2011)</xref> dan <xref ref-type="bibr" rid="BIBR-30">(Stewart et al., 2013)</xref> pada berbagai jenis nyamuk. </p><p>Dinotefuran termasuk insektisida baru dari kelompok <italic>furanicotinyl</italic> generasi ketiga dari kelompok neonikotinoid. Dinotefuran merupakan insektisida racun kontak dan racun perut yang bertindak sebagai agonis reseptor <italic>nicotinic acetylcholine</italic>. Dinotefuran juga bersifat non-repelen, sedikit iritan dan toksisitas rendah terhadap mamalia <xref ref-type="bibr" rid="BIBR-19">(Chemical, 2021)</xref><xref ref-type="bibr" rid="BIBR-12">(Committee, 2021)</xref>. Penggunaan dinotefuran dalam aplikasi ATSB dilaporkan <xref ref-type="bibr" rid="BIBR-14">(Khallayoune et al., 2013)</xref> dan <xref ref-type="bibr" rid="BIBR-10">(Gu et al., 2020)</xref> terhadap beberapa jenis nyamuk. </p><p>Permetrin merupakan insektisida yang termasuk golongan sintetik piretroid generasi ketiga (1973). Insektisida sintetik piretroid bekerja pada gerbang sodium channel protein. Permetrin termasuk piretroid tipe I yang menyebabkan pelepasan berulang (repetitive discharge) impuls saraf serangga, memblokir konduksi dan memiliki koefisien suhu negatif. Permetrin mempunyai daya kontak cepat, daya bunuh tinggi (killing agent), toksisitas mamalia rendah, daya residu rendah hingga sedang, dan bersifat repelen <xref ref-type="bibr" rid="BIBR-32">(Wirawan, 2006)</xref><xref ref-type="bibr" rid="BIBR-12">(Committee, 2021)</xref>. Penggunaan permetrin dalam aplikasi ATSB dilaporkan <xref ref-type="bibr" rid="BIBR-3">(Allan, 2011)</xref> dan <xref ref-type="bibr" rid="BIBR-27">(Shin et al., 2011)</xref> terhadap berbagai jenis nyamuk.</p><p>Deltametrin merupakan insektisida yang termasuk golongan sintetik piretroid generasi keempat. Deltametrin termasuk piretroid tipe II yang memiliki koefisien suhu positif dan bekerja dengan tidak menyebabkan pelepasan berulang (repetitive discharge) impuls saraf, seperti piretroid tipe I <xref ref-type="bibr" rid="BIBR-32">(Wirawan, 2006)</xref><xref ref-type="bibr" rid="BIBR-12">(Committee, 2021)</xref>. Penggunaan deltametrin dalam aplikasi ATSB dilaporkan oleh <xref ref-type="bibr" rid="BIBR-3">(Allan, 2011)</xref>, <xref ref-type="bibr" rid="BIBR-27">(Shin et al., 2011)</xref>, dan <xref ref-type="bibr" rid="BIBR-10">(Gu et al., 2020)</xref> terhadap berbagai jenis nyamuk.</p><p>Hasil penelitian ini merupakan informasi baru yang sangat bernilai terutama dalam mengendalikan lipas dengan cara memanfaatkan ketertarikan lipas terhadap gula yang kemudian dibubuhkan bahan toksik sehingga menimbulkan kematian lipas. Bahan toksik yang dapat digunakan berupa insektisida yang saat ini mudah didapatkan di pasar. Insektisida yang beredar di Indonesia terdiri atas berbagai golongan, seperti piretroid, karbamat, organofosfat, neonikotonoid, pirol, dan lain-lain. Aplikasi ATSB dapat dilakukan dengan mudah dengan menggunakan berbagai wadah yang dapat disesuaikan dengan situasi setempat, kemudian ditempatkan pada lokasi yang sering ditemukan lipas. Hasil ini juga menunjukkan keefektifan yang berbeda-beda, sebagai contoh meskipun sama-sama satu golongan piretroid, deltametrin lebih efektif (angka kematian 24 jam sebesar 100%) dibandingkan dengan permetrin dengan angka kematian 52,5% pada waktu pengamatan yang sama. Sesungguhnya setiap golongan insektisida mempunyai sifat dan <italic>mode of action</italic> yang berda-beda sehingga keefektifannya hanya bisa dibandingkan dengan sesama insektisida dengan golongan, konsentrasi, dan pengamatan yang sama.</p></sec><sec><title>KESIMPULAN</title><p>Konsentrasi umpan gula 5% merupakan bahan paling menarik untuk didatangi oleh lipas jerman <italic>B. germanica</italic>. Bahan ATSB berupa umpan gula 5% dengan bahan toksik dari asam borat, klorfenapir, dinotefuran, permetrin, dan deltametrin efektif membunuh lipas <italic>B. germanica</italic> dengan tingkat kematian yang bervariasi dalam waktu 24 jam pascaperlakuan. ATSB dengan bahan toksik dinetofuran dan deltametrin efektif menimbulkan kematian lipas tertinggi (100%). 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